Mark Lipson et al.
Parallel ancient genomic transects reveal complex population history of early European farmersdoi:
https://doi.org/10.1101/114488 Abstract
Ancient DNA studies have established that European Neolithic populations were descended from Anatolian migrants who received a limited amount of admixture from resident hunter-gatherers. Many open questions remain, however, about the spatial and temporal dynamics of population interactions and admixture during the Neolithic period. Using the highest-resolution genome-wide ancient DNA data set assembled to date---a total of 177 samples, 127 newly reported here, from the Neolithic and Chalcolithic of Hungary (6000-2900 BCE, n = 98), Germany (5500-3000 BCE, n = 42), and Spain (5500-2200 BCE, n = 37)---we investigate the population dynamics of Neolithization across Europe. We find that genetic diversity was shaped predominantly by local processes, with varied sources and proportions of hunter-gatherer ancestry among the three regions and through time. Admixture between groups with different ancestry profiles was pervasive and resulted in observable population transformation across almost all cultural transitions. Our results shed new light on the ways that gene flow reshaped European populations throughout the Neolithic period and demonstrate the potential of time-series-based sampling and modeling approaches to elucidate multiple dimensions of historical population interactions.
Sample ID Lab ID Skeletal element #Libraries Reference Date Culture Location Country Lat. Long. Sex mtDNA haplogroup Y chromosome haplogoup
GEN68 I2794 Petrous (cohlea, left) 1 Unpublished 5706-5541 calBCE (6700±40 BP, Poz-83628) Koros_EN Törökszentmiklós, road 4, site 3 Hungary 47,191612 20,402331 F K1a26 ..
HUNG276, KO2 I1508 Petrous 1 MathiesonNature2015 (capture of same sample shotgunned in Gamba2014) 5713-5566 calBCE (6726±35 BP, OxA-28101) Koros_EN Tiszaszőlős-Domaháza Hungary 47,318741 21,527443 F K1a ..
BAM17b I1878 Petrous 1 Unpublished 5832-5667 calBCE (6857±31 BP, MAMS-11935) Starcevo_EN Alsónyék-Bátaszék, Mérnöki telep Hungary 46,2055 18,70503 M T1a2 H2
BAM25 I0174 Tooth (M46/M47) 1 MathiesonNature2015 (1240k of same same sample with 390k in HaakLazaridis2015) 5702-5536 calBCE (6695±40BP, MAMS-11939 ) Starcevo_EN Alsónyék-Bátaszék, Mérnöki telep Hungary 46,208562 18,700047 M N1a1a1 H2
BAM4a I1876 Tooth (M36) 1 Unpublished 5641-5547 calBCE (6677±27 BP, MAMS-11928) Starcevo_EN Alsónyék-Bátaszék, Mérnöki telep Hungary 46,2055 18,70503 M K1a4 G2a2a1
LGCS1a I1880 Tooth (M17) 1 Unpublished 5800-5500 BCE Starcevo_EN Lánycsók, Gata-Csatola Hungary 45,993 18,58127 M W5 G2a2b2b1a
BAL25b I1904 Petrous (right) 1 Unpublished 5208-4948 calBCE (6115±35 BP, Poz-82584) LBKT_MN Bátaszék-Lajvér Hungary 46,20487 18,7004 M K1b1a1 G2a2a1
BOVO1b I4181 Tooth (premolar) 1 Unpublished 5300-4900 BCE LBKT_MN Bölcske-Gyűrűsvölgy Hungary 46,736119 18,958091 F H27 ..
BUD4a I1882 Tooth (M26) 1 Unpublished 5300-4900 BCE LBKT_MN Budakeszi, Szőlőskert-Tangazdaság Hungary 47,50163 18,910468 M T1a G2a2b2a
BUD9a I4196 Tooth (M36) 1 Unpublished 5300-4900 BCE LBKT_MN Budakeszi, Szőlőskert-Tangazdaság Hungary 47,50163 18,91047 F U2 ..
GEN18 I2739 Petrous (cohlea, right) 1 Unpublished 5309-5074 calBCE (6244±34 BP, SUERC-51459) LBKT_MN Alsónyék-Bátaszék, site 11 Hungary 46,208562 18,700047 M T2c G2a2b2b1
KON3 I3536 Tooth (M18) 1 Unpublished 5300-4900 BCE LBKT_MN Enese elkerülő, Kóny, Proletár-dülö, M85, Site 2 Hungary 47,63868 17,36453 F T2b ..
SZEH4 I0176 Tooth (M24) 1 MathiesonNature2015 (1240k of same same sample with 390k in HaakLazaridis2015) 5207-4944 calBCE (6110±30 BP, Beta-310038) LBKT_MN Szemely-Hegyes Hungary 46,4 18,74 F N1a1a1a3 ..
CEG07b I2744 Tooth (M37) 1 Unpublished 5300-4900 BCE ALPc_Szakalhat_MN Cegléd, site 4/2 Hungary 47,178841 19,860515 M J2b1 G2a2b2a
CEG08b I2745 Tooth (m75) 1 Unpublished 5300-4900 BCE ALPc_Szakalhat_MN Cegléd, site 4/3 Hungary 47,178841 19,860515 F J1c1 ..
EBSA2a I4186 Tooth (M26) 1 Unpublished 5300-4900 BCE ALPc_Esztar_MN Ebes-Sajtgyár Hungary 47,477645 21,503001 F K1a ..
EBVO5a I4187 Tooth (M26) 1 Unpublished 5300-4900 BCE ALPc_Esztar_MN Ebes-Zsongvölgy Hungary 47,480426 21,501858 M V1a CT
HAJE10a I3535 Tooth (M17) 1 Unpublished 5221-5000 calBCE (6170±40 BP, Poz-83632) ALPc_III_MN Hajdúnánás-Eszlári út Hungary 47,85755 21,43239 M J2b1 I
HAJE7a I2384 Tooth (M47) 1 Unpublished 5302-5057 calBCE (6220±40 BP, Poz-83631) ALPc_III_MN Hajdúnánás-Eszlári út Hungary 47,85755 21,43239 M K1a26 I2
HELI11a I2379 Tooth (M16/26) 1 Unpublished 5209-4912 cal BCE (6100 ± 40 BP, Poz-88115) ALPc_Tiszadob_MN Hejőkürt-Lidl logisztikai központ Hungary 47,86071 21,00058 M N1a1 I2a2a1b
HELI2a I2378 Tooth (M28) 1 Unpublished 5300-4900 BCE ALPc_Tiszadob_MN Hejőkürt-Lidl logisztikai központ Hungary 47,86071 21,00058 M U8b1b I
HUNG302, NE2 I1498 Petrous 1 MathiesonNature2015 (capture of same sample shotgunned in Gamba2014) 5291-5056 calBCE (6207±30 BP, OxA-27858) ALPc_MN Debrecen Tócópart Erdőalja Hungary 47,52 21,589 F H ..
HUNG372, NE5 I1500 Petrous 1 MathiesonNature2015 (capture of same sample shotgunned in Gamba2014) 5295-4950 calBCE (6164±64 BP,OxA-23763) ALPc_MN Kompolt-Kígyósér Hungary 47,167 20,833 M J1c1 C1a2
HUNG86, NE3 I1499 Petrous 1 MathiesonNature2015 (capture of same sample shotgunned in Gamba2014) 5281-5026 calBCE (6185±34 BP, OxA-27732) Bükk_MN Garadna-Elkerülő út site 2 Hungary 48,52 21,168 F X2b-T226C ..
MEMO24b I2382 Tooth (M27) 1 Unpublished 5500-5300 BCE ALPc_Szatmar_MN Mezőkövesd-Mocsolyás Hungary 47,78134 20,58258 M U8b1b CT
MEMO2b I2380 Tooth (PM) 1 Unpublished 5500-5300 BCE ALPc_Szatmar_MN Mezőkövesd-Mocsolyás Hungary 47,78134 20,58258 F K1a1 ..
MEMO7a I3537 Tooth (M26) 1 Unpublished 5500-5300 BCE ALPc_MN Mezőkövesd-Mocsolyás Hungary 47,78134 20,58258 F HV ..
PF325, NE1 I1506 Petrous 1 MathiesonNature2015 (capture of same sample shotgunned in Gamba2014) 5306-5071 calBCE (6237±32 BP, OxA-27861) ALPc_MN Polgár-Ferenci-hát Hungary 47,88 21,192 F U5b2c ..
PF839/1198, NE4 I1505 Petrous 1 MathiesonNature2015 (capture of same sample shotgunned in Gamba2014) 5211-5011 calBCE (6153±33 BP, OxA-28020) ALPc_MN Polgár-Ferenci-hát Hungary 47,88 21,192 F J1c5 ..
POPI5a I4188 Tooth (M36) 1 Unpublished 5300-4900 BCE ALPc_I_MN Polgár-Piócás Hungary 47,865071 21,125287 M K1a1 I2a2a
PULE1.18a I2355 Tooth (M46) 1 Unpublished 5300-4900 BCE ALPc_Szakalhat_MN Pusztataskony-Ledence I. Hungary 47,45925 20,51282 F T2c2 ..
PULE1.23a I2357 Tooth (M17) 1 Unpublished 5300-4900 BCE ALPc_Szakalhat_MN Pusztataskony-Ledence I. Hungary 47,45925 20,51282 F H1e ..
TISO13a I2377 Tooth (M46) 1 Unpublished 5208-4942 calBCE (6110±40 BP, Poz-83630) ALPc_Tiszadob_Bükk_MN Tiszadob-Ó-Kenéz Hungary 48,00363 21,17651 M J1c2 I2a2a
TISO1b I2375 Tooth (C33/43) 1 Unpublished 5300-4900 BCE ALPc_Tiszadob_Bükk_MN Tiszadob-Ó-Kenéz Hungary 48,00363 21,17651 M H7 I2a2a1b1
TISO3a I4199 Tooth (M16) 1 Unpublished 5300-4900 BCE ALPc_Tiszadob_Bükk_MN Tiszadob-Ó-Kenéz Hungary 48,00363 21,17651 F U5b2b1a ..
SEKU10a I1896 Tooth (M16) 1 Unpublished 5320-5080 calBCE (6264±34 BP) Vinca_MN Szederkény-Kukorica-dülö Hungary 45,5969 18,28247 M K2a G2a2b2a1a
SEKU6a I1895 Tooth (M26) 1 Unpublished 5321-5081 calBCE (6267±33 BP) Vinca_MN Szederkény-Kukorica-dülö Hungary 45,5969 18,28247 F H26 ..
VEGI17a I1894 Tooth (M48) 1 Unpublished 5400-5000 BCE Vinca_MN Versend-Gilencsa Hungary 45,5853 18,30771 F U2 ..
VEGI3a I1887 Tooth (M46) 1 Unpublished 5400-5000 BCE Vinca_MN Versend-Gilencsa Hungary 45,5853 18,30771 M T2b H2
Gorzsa18 I0449 .. 1 Unpublished 5000-4500 BCE Tisza_LN Hódmezővásárhely-Gorzsa grave 18 Hungary 46,36 20,43 M U5b2c I2a1
Gorzsa4 I0447 .. 1 Unpublished 5000-4500 BCE Tisza_LN Hódmezővásárhely-Gorzsa grave 4 Hungary 46,36 20,43 F T1a ..
KOKE3a I2387 Tooth (M46) 1 Unpublished 5000-4500 BCE Tisza_LN Hódmezővásárhely-Kökénydomb Vörös tanya Hungary 46,394 20,24495 M K1b1a I
PULE1.24 I2358 Tooth (M16) 1 Unpublished 5000-4500 BCE Tisza_LN Pusztataskony-Ledence I. Hungary 47,45925 20,51282 F K1a4f ..
VSM3a I2746 Tooth (M17) 1 Unpublished 5000-4500 BCE Tisza_LN Vésztő-Mágor Hungary 46,939581 21,20907 M H26 G2a
ALE14a I1893 Tooth (m85) 2 Unpublished 5030-4848 calBCE (6049±29 BP, MAMS-14817) Sopot_LN Alsónyék-Elkerülő 2. site Hungary 46,2041 18,7225 M U8b1 G2a
ALE4a i4185 Tooth (M36) 1 Unpublished 5016-4838 calBCE (6032±32 BP, MAMS-14814) Sopot_LN Alsónyék-Elkerülő 2. site Hungary 46,204523 18,729916 M T2c F
BAL3a I1903 Tooth (M46) 1 Unpublished 4800-4500 BCE Lengyel_LN Bátaszék-Lajvér Hungary 46,20487 18,7004 M T2f H1b1
CSAT19a I1906 Tooth (M46) 1 Unpublished 4800-4500 BCE Lengyel_LN Csabdi-Télizöldes Hungary 47,51062 18,61693 M H H
CSAT25a I1905 Petrous (cohlea, left) 1 Unpublished 4826-4602 calBCE (5850±40 BP, Poz-82582) Lengyel_LN Csabdi-Télizöldes Hungary 47,51062 18,61693 M T2b I2
FAGA1a I1890 Tooth (M36) 1 Unpublished 5100-4750 BCE Sopot_LN Fajsz-Garadomb Hungary 46,41594 18,91974 M HV0a I
FAGA2a I1891 Tooth (M46) 1 Unpublished 5195-4842 calBCE (6060±40 BP, Poz-83633) Sopot_LN Fajsz-Garadomb Hungary 46,41594 18,91974 F H ..
FEB3a I1902 Tooth (PM44) 1 Unpublished 4800-4500 BCE Lengyel_LN Felsőörs-Bárókert Hungary 47,01938 17,96243 M H44b J2a
HUNG347, NE7 I1495 Petrous 1 MathiesonNature2015 (capture of same sample shotgunned in Gamba2014) 4491-4357 calBCE (5598±32 BP, MAMS-14819) Lengyel_LN Apc-Berekalja Hungary 47,167 19,833 M N1a1a1a I
SZEH5a i4183 Tooth (M26) 1 Unpublished 4904-4709 calBCE (5920±40 BP, Beta-310039) Sopot_LN Szemely-Hegyes Hungary 46,025398 18,325056 M K1b1a G
SZEH7b i4184 Petrous 1 Unpublished 4930-4715 calBCE (5930±40 BP, Beta-310040) Sopot_LN Szemely-Hegyes Hungary 46,025398 18,325056 F K1a26 ..
VEJ12a I2352 Tooth (M16) 1 Unpublished 4800-4500 BCE Lengyel_LN Veszprém Jutasi út Hungary 47,102645 17,912987 M U8b1a H
VEJ2a I1899 Tooth (M38) 1 Unpublished 4800-4500 BCE Lengyel_LN Veszprém Jutasi út Hungary 47,102645 17,912987 M T2b C
VEJ5a I1901 Tooth (M38) 1 Unpublished 4800-4500 BCE Lengyel_LN Veszprém Jutasi út Hungary 47,102645 17,912987 M J1c2 G2a2a1
GEN67 I2793 Petrous (cohlea, right) 1 Unpublished 4444-4257 calBCE (5480±35 BP, Poz-83629) Tiszapolgar_Bodrogkeresztur_ECA Törökszentmiklós, road 4, site 3 Hungary 47,191612 20,402331 M H1a I2a2a1b
PULE1.10a I2353 Tooth (M17) 1 Unpublished 4500-4000 BCE Tiszapolgar_ECA Pusztataskony-Ledence I. Hungary 47,45925 20,51282 M T2c I2a
PULE1.13a I2354 Tooth (M37) 1 Unpublished 4500-4000 BCE Tiszapolgar_ECA Pusztataskony-Ledence I. Hungary 47,45925 20,51282 M T2c G2a2b2a1a1c1a
PULE1.9a I2395 Tooth (M48) 1 Unpublished 4500-4000 BCE Tiszapolgar_ECA Pusztataskony-Ledence I. Hungary 47,45925 20,51282 M H26 G2a2b
GEN100 i4189 Petrous (cohlea, left) 1 Unpublished 4300-3900 BCE Balaton_Lasinja_MCA Alsónyék, site 11 Hungary 46,204523 18,729916 F T2b ..
GEN49 I2783 Petrous (cohlea, left) 1 Unpublished 4228-3963 calBCE (5230±40 BP, Poz-83638) Hunyadihalom_MCA Nemesnádudvar-Papföld, M9/7 lh. Hungary 46,334774 19,053945 M T2b23 CT
KEFP2a I1908 Petrous 1 Unpublished 4300-3900 BCE Balaton_Lasinja_MCA Keszthely-Fenékpuszta, Pusztaszentegyházi-dűlő Hungary 46,70991 17,23935 F J2b1a ..
KON2a I1907 Tooth (M36) 1 Unpublished 4333-4072 calBCE (5380±30 BP, Beta-310033) Balaton_Lasinja_MCA Enese elkerülő, Kóny, Proletár-dülö, M85, Site 2 Hungary 47,63868 17,36453 F K2a ..
M6-116.12a I1909 Tooth (M37) 1 Unpublished 4232-4046 calBCE (5300±23 BP, MAMS-14132) Balaton_Lasinja_MCA Lánycsók, Csata-alja Hungary 45,99604 18,58101 F T2f ..
VEJ9a I2394 Tooth (M46) 1 Unpublished 4339-4237 calBCE (5418±29 BP, MAMS-14828) Balaton_Lasinja_MCA Veszprém Jutasi út Hungary 47,102645 17,912987 M H40 CT
GEN60 I2788 Petrous (cohlea, left) 1 Unpublished 3909-3651 calBCE (4960±40 BP, VERA-5402) Protoboleraz_LCA Abony, Turjányos-dűlő Hungary 47,189141 20,004795 M H G2a2b2a
GEN61 I2789 Petrous (cohlea, left) 1 Unpublished 3800-3600 BCE Protoboleraz_LCA Abony, Turjányos-dűlő Hungary 47,189141 20,004795 M J1c3b I2c
GEN62 I2790 Petrous (cohlea, left) 1 Unpublished 3800-3600 BCE Protoboleraz_LCA Abony, Turjányos-dűlő Hungary 47,189141 20,004795 F N1a1a3 ..
GEN63 I2791 Petrous (cohlea, left) 1 Unpublished 3762-3636 calBCE (4890±35 BP, VERA-5160) Protoboleraz_LCA Abony, Turjányos-dűlő Hungary 47,189141 20,004795 M U5a1c1 I2c
GEN12a I2366 Petrous (cohlea, left) 1 Unpublished 3300-2850 BCE Baden_LCA Budakalász-Luppa csárda Hungary 47,62094 19,04494 M H26a G2a2b2a1a1b1
GEN13a I2367 Petrous (cohlea, right) 1 Unpublished 3300-2850 BCE Baden_LCA Budakalász-Luppa csárda Hungary 47,62094 19,04494 M HV G2a2b2a1a
GEN15a I2369 Petrous (cohlea, left) 1 Unpublished 3367-3103 calBCE (4545±35 BP, Poz-83634) Baden_LCA Budakalász-Luppa csárda Hungary 47,62094 19,04494 M J2a1a1 G2a2b2a1a1c1a
GEN16a I2370 Petrous (cohlea, right) 1 Unpublished 3300-2850 BCE Baden_LCA Alsónémedi Hungary 47,31875 19,16692 F T2b ..
GEN17a I2371 Petrous (cohlea, right) 1 Unpublished 3359-3098 calBCE (4520±35 BP, Poz-83635) Baden_LCA Alsónémedi Hungary 47,31875 19,16692 M U5b3f G2a2a
GEN21 I2752 Petrous (cohlea, right) 1 Unpublished 3600-2850 BCE Baden_LCA Balatonlelle-Felső-Gamász Hungary 46,78469 17,731714 M K1a1 I2a1
GEN22 I2753 Petrous (cohlea, right) 1 Unpublished 3600-2850 BCE Baden_LCA Balatonlelle-Felső-Gamász Hungary 46,78469 17,731714 M U5a1 I2a1a1
GEN55 I2785 Petrous (cohlea, left) 1 Unpublished 3600-2850 BCE Baden_LCA Vámosgyörk MHAT telep Hungary 47,684976 19,918434 F T2c2 ..
HUNG353, CO1 I1497 Petrous 1 MathiesonNature2015 (capture of same sample shotgunned in Gamba2014) 3315-2923 calBCE (4421±27 BP, MAMS-14825) Baden_LCA Apc-Berekalja Hungary 47,167 19,833 F H ..
Vors1 I2763 Tooth (M28) 1 Unpublished 3300-2850 BCE Baden_LCA Vörs Hungary 46,665765 17,270774 F T2f ..
HAL03a I2005 Tooth 1 Unpublished 5295-5057 calBCE (6211±32 BP, KIA-40346) LBK_EN Halberstadt-Sonntagsfeld Germany 51,89583333 11,04666667 F T2b ..
HAL07a I2008 Tooth 1 Unpublished 5212-4992 calBCE (6137±35 BP, KIA-40342) LBK_EN Halberstadt-Sonntagsfeld Germany 51,89583333 11,04666667 F N1a1b ..
HAL15a I2014 .. 1 Unpublished 5199-4857 calBCE (KIA-40344; 6081±30 BP?) LBK_EN Halberstadt-Sonntagsfeld Germany 51,89583333 11,04666667 M N1a1a3 G2
HAL17b I2016 Bone 1 Unpublished 5500-4850 BCE LBK_EN Halberstadt-Sonntagsfeld Germany 51,89583333 11,04666667 F V1 ..
HAL18a I2017 Tooth 1 Unpublished 5500-4850 BCE LBK_EN Halberstadt-Sonntagsfeld Germany 51,89583333 11,04666667 F K2a ..
HAL19 I1550 Tooth 2 New libraries added to MathiesonNature2015 5500-4850 BCE LBK_EN Halberstadt-Sonntagsfeld Germany 51,89583333 11,04666667 F K1a2 ..
HAL2 I0659 Tooth 2 New libraries added in to previously published data (MathiesonNature2015, which itself was more capture on sample with original 390k data in HaakLazaridis2015) 5211-4963 calBCE (KIA-40350; 6130±40 BP?) LBK_EN Halberstadt-Sonntagsfeld Germany 51,89583333 11,04666667 M N1a1a1a2 G2a2a1
HAL20b I2020 Tooth 1 Unpublished 5500-4850 BCE LBK_EN Halberstadt-Sonntagsfeld Germany 51,89583333 11,04666667 M K1a2 G2a2a
HAL21a I2021 Tooth 1 Unpublished 5500-4850 BCE LBK_EN Halberstadt-Sonntagsfeld Germany 51,89583333 11,04666667 M T2b G2a2a
HAL22b I2022 Tooth 1 Unpublished 5500-4850 BCE LBK_EN Halberstadt-Sonntagsfeld Germany 51,89583333 11,04666667 F T2b ..
HAL24 I0821 Tooth 2 New libraries added in to previously published data (MathiesonNature2015, which itself was more capture on sample with original 390k data in HaakLazaridis2015) 5201-4850 calBCE (6076±34 BP, KIA-40348) LBK_EN Halberstadt-Sonntagsfeld Germany 51,89583333 11,04666667 M X2d1 G2a2a1
HAL25 I0048 Tooth 2 New libraries added in to previously published data (MathiesonNature2015, which itself was more capture on sample with original 390k data in HaakLazaridis2015) 5210-5002 calBCE (6153±33 BP, MAMS-21482) LBK_EN Halberstadt-Sonntagsfeld Germany 51,89583333 11,04666667 M K1a G2a2a1
HAL27a I2026 Bone 1 Unpublished 5500-4850 BCE LBK_EN Halberstadt-Sonntagsfeld Germany 51,89583333 11,04666667 M N1a1a3 G2a2a
HAL31a I2029 Tooth 1 Unpublished 5295-5057 calBCE (6211±32 BP, KIA-40349) LBK_EN Halberstadt-Sonntagsfeld Germany 51,89583333 11,04666667 F K1c ..
HAL32b I2030 Tooth 1 Unpublished 5500-4850 BCE LBK_EN Halberstadt-Sonntagsfeld Germany 51,89583333 11,04666667 F H26 ..
HAL34 I0057 Bone 2 New libraries added in to previously published data (MathiesonNature2015, which itself was more capture on sample with original 390k data in HaakLazaridis2015) 5219-5021 calBCE (6173±34 BP, MAMS-21483) LBK_EN Halberstadt-Sonntagsfeld Germany 51,89583333 11,04666667 F N1a1a1 ..
HAL35b I2032 Tooth 1 Unpublished 5500-4850 BCE LBK_EN Halberstadt-Sonntagsfeld Germany 51,89583333 11,04666667 F J1c ..
HAL38a I2036 Tooth 1 Unpublished 5500-4850 BCE LBK_EN Halberstadt-Sonntagsfeld Germany 51,89583333 11,04666667 F V1 ..
HAL39b I2037 Tooth 1 Unpublished 5210-5002 calBCE (6144±32 BP, KIA-40343) LBK_EN Halberstadt-Sonntagsfeld Germany 51,89583333 11,04666667 M H1e G2a2a1
HAL4 I0100 Tooth 3 New libraries added in to previously published data (MathiesonNature2015, which itself was more capture on sample with original 390k data in HaakLazaridis2015) 5202-4852 calBCE (6080±32 BP, KIA-40341) LBK_EN Halberstadt-Sonntagsfeld Germany 51,89583333 11,04666667 F N1a1a1a ..
HAL40a I2038 Bone 1 Unpublished 5500-4850 BCE LBK_EN Halberstadt-Sonntagsfeld Germany 51,89583333 11,04666667 F T2b ..
HAL5 I0046 Tooth 5 New libraries added in to previously published data (MathiesonNature2015, which itself was more capture on sample with original 390k data in HaakLazaridis2015) 5211-4991 calBCE (MAMS-21479; 6136±34 BP?) LBK_EN Halberstadt-Sonntagsfeld Germany 51,89583333 11,04666667 F T2c1 ..
KAR16A I0797 Tooth 1 MathiesonNature2015 5500-4850 BCE LBK_EN Karsdorf Germany 51,28305556 11,65 M H46b T1a
KAR6 I0795 Tooth 1 MathiesonNature2015 (1240k of same same sample with 390k in HaakLazaridis2015) 5217-5041 calBCE (6174±29 BP, MAMS-22823) LBK_EN Karsdorf Germany 51,28305556 11,65 M H1/H1au1b CT
LBK1976 I0022 Tooth (right upper P2) 1 MathiesonNature2015 (1240k of same same sample with 390k in HaakLazaridis2015) 5500-4850 BCE LBK_EN Viesenhaeuser Hof, Stuttgart-Muehlhausen Germany 48,78 9,18 F T2e ..
LBK1992 I0025 Tooth (left lower M3) 1 MathiesonNature2015 (1240k of same same sample with 390k in HaakLazaridis2015) 5500-4850 BCE LBK_EN Viesenhaeuser Hof, Stuttgart-Muehlhausen Germany 48,78 9,18 F T2b ..
LBK2155 I0026 Tooth (left lower M3) 1 MathiesonNature2015 (1240k of same same sample with 390k in HaakLazaridis2015) 5500-4850 BCE LBK_EN Viesenhaeuser Hof, Stuttgart-Muehlhausen Germany 48,78 9,18 F T2b ..
Stuttgart I0018 Tooth 1 New libraries for previously published sample (LazaridisNature2014) 5310-5076 calBCE (6246±30 BP, MAMS-24635) LBK_EN Viesenhaeuser Hof, Stuttgart-Muehlhausen Germany 48,78 9,18 F T2c1b ..
UWS4 I0054 .. 2 MathiesonNature2015 (1240k of same same sample with 390k in HaakLazaridis2015) 5223-5021 calBCE (6180±34 BP, MAMS-21485) LBK_EN Unterwiederstedt Germany 51,66 11,53 F J1c17 ..
ESP30 I0807 Bone 1 MathiesonNature2015 (1240k of same same sample with 390k in HaakLazaridis2015) 3970-3710 calBCE (5061±62 BP, Er-7784) Baalberge_MN Esperstedt Germany 51,42 11,68 M H1e1a I
HAL13a I2012 .. 1 Unpublished 4600-4300 BCE Roessen_MN Halberstadt-Sonntagsfeld Germany 51,89583333 11,04666667 F V1a ..
QLB15D I0559 Tooth 1 MathiesonNature2015 (1240k of same same sample with 390k in HaakLazaridis2015) 3654-3527 calBCE (4815±26 BP, MAMS-22818) Baalberge_MN Quedlinburg, Site IX Germany 51,79 11,14 M HV R
QLB18A I0560 Tooth 1 MathiesonNature2015 (1240k of same same sample with 390k in HaakLazaridis2015) 3640-3376 calBCE (4745±52 BP, Er-7856) Baalberge_MN Quedlinburg, Site IX Germany 51,79 11,14 F T2e1 ..
SALZ3B I0551 Bone (tibia/Femur) 1 MathiesonNature2015 3400-3025 BCE Salzmuende_MN Salzmuende-Schiebzig Germany 51,53305556 11,83305556 M U3a1 G2a2a1
SALZ57A I0800 Tooth 1 Unpublished 3345-3097 calBCE (4498±27 BP, KIA-31459) Salzmuende_MN Salzmuende-Schiebzig Germany 51,53305556 11,83305556 F H3 ..
SALZ77A I0802 Tooth 1 Unpublished 3400-3025 BCE Salzmuende_MN Salzmuende-Schiebzig Germany 51,53305556 11,83305556 M H3 IJK (x J)
Bla16 I1593 Bone 3 Unpublished 3958-3344 calBCE [3512-3344 calBCE (4615±30, KIA-28845, Bla16); 3958-3773 calBCE (5055±35, KIA-37508, Bla27)] Blatterhohle_MN Blatterhole Cave Germany 51,358333 7,551111 M U5b2a2 R1b1
Bla28 I1594 Bone (mandible) 1 Unpublished 3337-3024 calBCE (4465±30, KIA-28846) Blatterhohle_MN Blatterhole Cave Germany 51,358333 7,551111 M J1c1b1 R1
Bla5 I1563 Bone 6 Unpublished 3704-3117 calBCE [3498-3117 calBCE (4580±30, KIA-28844, Bla5); 3704-3539 calBCE (4860±30, KIA-45011, Bla7); 3634-3378 calBCE (4730±25, KIA-45010, Bla13)] Blatterhohle_MN Blatterhole Cave Germany 51,358333 7,551111 F H5 ..
Bla8 I1565 Bone 7 Unpublished 4038-3532 calBCE [3786-3657 calBCE (4950±30, KIA-45006, Bla8); 3713-3642 calBCE (4905±25, KIA-45008, Bla9); 4038-3810 calBCE (5145±30, KIA-45007, Bla11); 3703-3532 calBCE (4845±35, KIA-37507, Bla24)] Blatterhohle_MN Blatterhole Cave Germany 51,358333 7,551111 M U5b2b2 I2a1
CB13 CB13.SG .. .. OlaldeMBE2015 5469-5327 calBCE (6410±30, Beta-384724) Iberia_EN.SG Cova Bonica, Vallirana, Barcelona Spain 41,37 1,89 F K1a2a ..
E-06-Ind1 I1972 Tooth 1 Unpublished 4827-4692 calBCE (5880±30 BP, Beta-366569) Iberia_EN El Prado de Pancorbo, Burgos Spain 42,6282 -3,11649 F K1a4a1 ..
E-14-Ind2 I2199 Tooth 1 Unpublished 5216-5031 calBCE (6170±30 BP, Beta-438208) Iberia_EN El Prado de Pancorbo, Burgos Spain 42,6282 -3,11649 F H1 ..
Troc1 I0409 Tooth 2 MathiesonNature2015 (1240k of same same sample with 390k in HaakLazaridis2015) 5311-5218 calBCE (6280±25 BP, MAMS-16159) Iberia_EN Els Trocs Spain 42,5 0,5 F J1c3 ..
Troc3 I0410 Tooth 1 MathiesonNature2015 (1240k of same same sample with 390k in HaakLazaridis2015) 5294-5066 calBCE (6217±25 BP, MAMS-16161) Iberia_EN Els Trocs Spain 42,5 0,5 M T2c1d/T2c1d2 R1b1a2
Troc5 I0412 Tooth 3 MathiesonNature2015 (1240k of same same sample with 390k in HaakLazaridis2015) 5310-5078 calBCE (6249±28 BP, MAMS-16164) Iberia_EN Els Trocs Spain 42,5 0,5 M N1a1a1 I2a1b1
Troc7 I0413 Tooth 1 MathiesonNature2015 (1240k of same same sample with 390k in HaakLazaridis2015) 5303-5075 calBCE (6234±28 BP, MAMS-16166) Iberia_EN Els Trocs Spain 42,5 0,5 F V ..
Mina18 I0408 Tooth 3 MathiesonNature2015 (1240k of same same sample with 390k in HaakLazaridis2015) 3893-3661 calBCE (4970±30 BP, Beta 316132) Iberia_MN La Mina Spain 41,2522 -2,32622 F U5b1 ..
Mina3 I0405 Tooth 1 MathiesonNature2015 (1240k of same same sample with 390k in HaakLazaridis2015) 3900-3600 BCE Iberia_MN La Mina Spain 41,2522 -2,32622 M K1a1b1 H2
Mina4 I0406 Tooth 1 MathiesonNature2015 (1240k of same same sample with 390k in HaakLazaridis2015) 3900-3600 BCE Iberia_MN La Mina Spain 41,2522 -2,32622 M H1 I2a2a1b2
Mina6 I0407 Tooth 1 MathiesonNature2015 (1240k of same same sample with 390k in HaakLazaridis2015) 3900-3600 BCE Iberia_MN La Mina Spain 41,2522 -2,32622 F K1b1a1 ..
1.-K11 I3272 Tooth 1 Unpublished 3263-2903 calBCE (4380±40 BP, Beta-288933) Iberia_CA La Chabola de la Hechicera, Alava, Basque country Spain 42,57 -2,55 M X2b I2a2
3.-K11 I3273 Tooth 1 Unpublished 3627-3363 calBCE (4670±40 BP, Beta-288935) Iberia_CA La Chabola de la Hechicera, Alava, Basque country Spain 42,57 -2,55 F J2a1a1 ..
5.-K18 I1975 Tooth 1 Unpublished 3090-2894 calBCE (4350±40 BP, Beta-288937) Iberia_CA La Chabola de la Hechicera, Alava, Basque country Spain 42,57 -2,55 M J1c1 I2a2
ES.1/4 I1976 Tooth 1 Unpublished 2571-2347 calBCE (3960±30 BP, Beta-299306) Iberia_CA El Sotillo, Alava, Basque Country Spain 42,57 -2,62 M H3 I
ES-6G-110 I2473 Tooth 1 Unpublished 2916-2714 calBCE (4250±30 BP, Beta-299305) Iberia_CA El Sotillo, Alava, Basque Country Spain 42,57 -2,62 M H3 I2a2a
Inventario0/4 I2467 Tooth 1 Unpublished 2481-2212 calBCE (3900±40 BP, Beta-299300) Iberia_CA El Sotillo, Alava, Basque Country Spain 42,57 -2,62 M X2b I2a2a
LHUE11J.5 I1843 Tooth 1 Unpublished 3092-2877 calBCE [3092-2918 calBCE (4390±30 BP, Beta-301225), 3011-2877 calBCE (4290±30 BP, Beta-301226)] Iberia_CA Alto de la Huesera, Alava, Basque country Spain 42,57 -2,57 F U5b1 ..
LHUE2010.10 I1981 Tooth 1 Unpublished 3014-2891 calBCE (4320±30 BP, Beta-301223) Iberia_CA Alto de la Huesera, Alava, Basque country Spain 42,57 -2,57 F J1c1 ..
LHUE2010.11 I3276 Tooth 1 Unpublished 3092-2918 calBCE (4390±30 BP, Beta-301225) Iberia_CA Alto de la Huesera, Alava, Basque country Spain 42,57 -2,57 M V G2a2a
LHUE2014.11J I3277 Tooth 1 Unpublished 3100-2850 BCE (based on other dated samples in same layer) Iberia_CA Alto de la Huesera, Alava, Basque country Spain 42,57 -2,57 F U5b2b ..
LY.II.A.10.15066 I3269 Tooth 1 Unpublished 3350-2750 BCE [3022-2779 calBCE (4290±40 BP, Beta-137895); 3090-2900 calBCE (4360±40 BP, Beta-137896); 3310-2904 calBCE (4390±40 BP, Beta-148054) three dates of the whole stratigraphy of the site] Iberia_CA Las Yurdinas II, Alava, Basque Country Spain 42,63 -2,7 M U5b2b3a I2a2a
LY.II.A.10.15067 I3270 Tooth 1 Unpublished 3350-2750 BCE [3022-2779 calBCE (4290±40 BP, Beta-137895); 3090-2900 calBCE (4360±40 BP, Beta-137896); 3310-2904 calBCE (4390±40 BP, Beta-148054) three dates of the whole stratigraphy of the site] Iberia_CA Las Yurdinas II, Alava, Basque Country Spain 42,63 -2,7 F J2a1a1 ..
LY.II.A.10.15068 I3271 Tooth 1 Unpublished 3350-2750 BCE [3022-2779 calBCE (4290±40 BP, Beta-137895); 3090-2900 calBCE (4360±40 BP, Beta-137896); 3310-2904 calBCE (4390±40 BP, Beta-148054) three dates of the whole stratigraphy of the site] Iberia_CA Las Yurdinas II, Alava, Basque Country Spain 42,63 -2,7 F K1a4a1a ..
LY.II.A.10.15069 I1838 Tooth 1 Unpublished 3354-2943 calBCE (4470±50 BP, Poz-83429) Iberia_CA Las Yurdinas II, Alava, Basque Country Spain 42,63 -2,7 F J1c3 ..
MIR1 I1271 Tooth (root) 1 MathiesonNature2015 2900-2346 BCE [2568-2346 calBCE (3950±30 BP, Beta-416457); 2857-2496 calBCE (4080±30 BP, Beta-416456); 2865-2575 calBCE (4110±30 BP, Beta-416458); 2900-2679 calBCE (4210±30 BP, Beta-416455); based on dates of other El Mirador samples) Iberia_CA El Mirador Cave, Atapuerca, Burgos Spain 42,33333333 -3,5 F K1a ..
MIR13 I1276 Tooth (root) 1 MathiesonNature2015 2900-2346 BCE [2568-2346 calBCE (3950±30 BP, Beta-416457); 2857-2496 calBCE (4080±30 BP, Beta-416456); 2865-2575 calBCE (4110±30 BP, Beta-416458); 2900-2679 calBCE (4210±30 BP, Beta-416455); based on dates of other El Mirador samples) Iberia_CA El Mirador Cave, Atapuerca, Burgos Spain 42,33333333 -3,5 F H3c3 ..
MIR14 I1277 Tooth (root) 1 MathiesonNature2015 2568-2346 calBCE (3950±30 BP, Beta-416457) Iberia_CA El Mirador Cave, Atapuerca, Burgos Spain 42,33333333 -3,5 M H3 I2a2a
MIR17 I1280 Tooth (root) 1 MathiesonNature2015 2900-2346 BCE [2568-2346 calBCE (3950±30 BP, Beta-416457); 2857-2496 calBCE (4080±30 BP, Beta-416456); 2865-2575 calBCE (4110±30 BP, Beta-416458); 2900-2679 calBCE (4210±30 BP, Beta-416455); based on dates of other El Mirador samples) Iberia_CA El Mirador Cave, Atapuerca, Burgos Spain 42,33333333 -3,5 F J1c1 ..
MIR18 I1281 Tooth (root) 1 MathiesonNature2015 2865-2575 calBCE (4110±30 BP, Beta-416458) Iberia_CA El Mirador Cave, Atapuerca, Burgos Spain 42,33333333 -3,5 F H1t ..
MIR19 I1282 Tooth (root) 1 MathiesonNature2015 2900-2346 BCE [2568-2346 calBCE (3950±30 BP, Beta-416457); 2857-2496 calBCE (4080±30 BP, Beta-416456); 2865-2575 calBCE (4110±30 BP, Beta-416458); 2900-2679 calBCE (4210±30 BP, Beta-416455); based on dates of other El Mirador samples) Iberia_CA El Mirador Cave, Atapuerca, Burgos Spain 42,33333333 -3,5 M H3 I
MIR2 I1272 Tooth (root) 1 MathiesonNature2015 2857-2496 calBCE (4080±30 BP, Beta-416456) Iberia_CA El Mirador Cave, Atapuerca, Burgos Spain 42,33333333 -3,5 F K1b1a1 ..
MIR21 I1284 Tooth (root) 1 MathiesonNature2015 2900-2346 BCE [2568-2346 calBCE (3950±30 BP, Beta-416457); 2857-2496 calBCE (4080±30 BP, Beta-416456); 2865-2575 calBCE (4110±30 BP, Beta-416458); 2900-2679 calBCE (4210±30 BP, Beta-416455); based on dates of other El Mirador samples) Iberia_CA El Mirador Cave, Atapuerca, Burgos Spain 42,33333333 -3,5 M H3 I
MIR22 I1300 Tooth (root) 1 MathiesonNature2015 2900-2346 BCE [2568-2346 calBCE (3950±30 BP, Beta-416457); 2857-2496 calBCE (4080±30 BP, Beta-416456); 2865-2575 calBCE (4110±30 BP, Beta-416458); 2900-2679 calBCE (4210±30 BP, Beta-416455); based on dates of other El Mirador samples) Iberia_CA El Mirador Cave, Atapuerca, Burgos Spain 42,33333333 -3,5 F K1a2a ..
MIR24 I1302 Tooth (root) 1 MathiesonNature2015 2900-2346 BCE [2568-2346 calBCE (3950±30 BP, Beta-416457); 2857-2496 calBCE (4080±30 BP, Beta-416456); 2865-2575 calBCE (4110±30 BP, Beta-416458); 2900-2679 calBCE (4210±30 BP, Beta-416455); based on dates of other El Mirador samples) Iberia_CA El Mirador Cave, Atapuerca, Burgos Spain 42,33333333 -3,5 M J2b1a3 G2a2b2b
MIR25 I1303 Tooth (root) 1 MathiesonNature2015 2900-2346 BCE [2568-2346 calBCE (3950±30 BP, Beta-416457); 2857-2496 calBCE (4080±30 BP, Beta-416456); 2865-2575 calBCE (4110±30 BP, Beta-416458); 2900-2679 calBCE (4210±30 BP, Beta-416455); based on dates of other El Mirador samples) Iberia_CA El Mirador Cave, Atapuerca, Burgos Spain 42,33333333 -3,5 M U3a1 I2a1a1
MIR5, MIR6 I0581 Tooth (root) 2 MathiesonNature2015 2900-2679 calBCE (4210±30 BP, Beta-416455) Iberia_CA El Mirador Cave, Atapuerca, Burgos Spain 42,33333333 -3,5 M X2b C1a2
Not being used for analyses:
BAM13b I1877 Petrous 1 Unpublished 5704-5556 calBCE (6704±34 BP, MAMS-11933) Starcevo_EN Alsónyék-Bátaszék, Mérnöki telep Hungary 46,2055 18,70503 M X2d1 G2a2b2b
CEG03b I2743 Tooth (M38) 1 Unpublished 5300-4900 BCE ALPc_Szakalhat_MN Cegléd, site 4/1 Hungary 47,178841 19,860515 M H G2a2b2a
GEN14a I2368 Petrous (cohlea, right) 1 Unpublished 3300-2850 BCE Baden_LCA Budakalász-Luppa csárda Hungary 47,62094 19,04494 M U5b2a2c G2a2b2a
GEN23 I2754 Petrous (cohlea, left) 1 Unpublished 3337-3024 calBCE (4465±30 BP, Poz-83637) Baden_LCA Balatonlelle-Felső-Gamász Hungary 46,78469 17,731714 M J1c I2
GEN24 I2755 Petrous (cohlea, right) 1 Unpublished 3600-2850 BCE Baden_LCA Balatonlelle-Felső-Gamász Hungary 46,78469 17,731714 M U8b1a1 I
HAJE1a I2383 Tooth (M28) 1 Unpublished 5300-4900 BCE ALPc_III_MN Hajdúnánás-Eszlári út Hungary 47,85755 21,43239 M T2b23 G2
HAL14 I0056 Tooth 2 New libraries added in to previously published data (MathiesonNature2015, which itself was more capture on sample with original 390k data in HaakLazaridis2015) 5213-5009 calBCE (6156±35 BP, MAMS-21480) LBK_EN Halberstadt-Sonntagsfeld Germany 51,89583333 11,04666667 M T2b G2a2a
PULE1.22a I2356 Tooth (M37) 1 Unpublished 4500-4000 BCE Tiszapolgar_ECA Pusztataskony-Ledence I. Hungary 47,45925 20,51282 M T2c G2a2b
PULE1.26a I2359 Tooth (M28) 1 Unpublished 5000-4500 BCE Tisza_LN Pusztataskony-Ledence I. Hungary 47,45925 20,51282 F T2f ..
SCHM2a I1560_d Tooth 1 Unpublished 3780-3099 BCE [3359-3099 calBCE (4521±34 BP, KIA 2384); 3780-3647 calBCE (4930±36 BP, KIA 2388); layer dates oldest and youngest of eight dated individuals in the same collective grave] Germany_MN Erwitte-Schmerlecke Germany 51,6 8,25 M J2b1a I
SEKU1a I1889 Tooth (M46) 1 Unpublished 5202-4851 calBCE (6079±33 BP, MAMS-14808) Vinca_MN Szederkény-Kukorica-dülö Hungary 45,5969 18,28247 M K1a2 G2a2a
TISO11a I2376 Tooth (M18) 1 Unpublished 5500-5000 BCE ALPc_Tiszadob_Bükk_MN Tiszadob-Ókenéz Hungary 48,00363 21,17651 M HV0a H
TOLM2a I2351 Tooth (M46) 1 Unpublished 4300-3900 BCE Balaton_Lasinja_CA Tolna-Mözs TO3 Hungary 46,407091 18,742154 F T2c2 ..
TOLM4a I1883 Tooth (M36) 1 Unpublished 5301-5076 calBCE (6233±23 BP, MAMS-14145) LBKT_MN Tolna-Mözs TO26 Hungary 46,407091 18,742154 M K1a1a G2a
TOSM1a I2373 Bone (femur, right) 1 Unpublished 6000-5500 BCE Koros_EN Törökszentmiklos Tiszapüspöki Karanycs haromag 3. lh. Hungary 47,18283 20,3509 F H ..
TOSM3a I2374 Bone (femur, right) 1 Unpublished 6000-5500 BCE Koros_EN Törökszentmiklos Tiszapüspöki Karanycs haromag 3. lh. Hungary 47,18283 20,3509 M K1a4f G
VEGI1a I1885 Tooth (M17) 1 Unpublished 5359-5224 calBCE (6321±28 BP, MAMS-14830) Vinca_MN Versend-Gilencsa Hungary 45,5853 18,30771 F T2b ..
VEJ4a I1900 Tooth (M36) 1 Unpublished 4796-4685 calBCE (5861±26 BP, MAMS-14827) Lengyel_LN Veszprém Jutasi út Hungary 47,102645 17,912987 M W1 E1b1b1a1b1