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https://www.biorxiv.org/content/10.1101/2020.03.25.004606v1

The Genomic Formation of Human Populations in East Asia

View ORCID ProfileChuan-Chao Wang, Hui-Yuan Yeh, Alexander N Popov, Hu-Qin Zhang, Hirofumi Matsumura, Kendra Sirak, Olivia Cheronet, Alexey Kovalev, Nadin Rohland, Alexander M Kim, Rebecca Bernardos, Dashtseveg Tumen, Jing Zhao, Yi-Chang Liu, Jiun-Yu Liu, Matthew Mah, Swapan Mallick, Ke Wang, Zhao Zhang, Nicole Adamski, Nasreen Broomandkhoshbacht, Kimberly Callan, Brendan J Culleton, Laurie Eccles, Ann Marie Lawson, Megan Michel, Jonas Oppenheimer, Kristin Stewardson, Shaoqing Wen, Shi Yan, Fatma Zalzala, Richard Chuang, Ching-Jung Huang, Chung-Ching Shiung, Yuri G Nikitin, Andrei V Tabarev, Alexey A Tishkin, Song Lin, Zhou-Yong Sun, Xiao-Ming Wu, Tie-Lin Yang, Xi Hu, Liang Chen, Hua Du, Jamsranjav Bayarsaikhan, Enkhbayar Mijiddorj, Diimaajav Erdenebaatar, Tumur-Ochir Iderkhangai, Erdene Myagmar, Hideaki Kanzawa-Kiriyama, Msato Nishino, Ken-ichi Shinoda, Olga A Shubina, Jianxin Guo, Qiongying Deng, Longli Kang, Dawei Li, Dongna Li, Rong Lin, Wangwei Cai, Rukesh Shrestha, Ling-Xiang Wang, Lanhai Wei, Guangmao Xie, Hongbing Yao, Manfei Zhang, Guanglin He, Xiaomin Yang, Rong Hu, Martine Robbeets, Stephan Schiffels, Douglas J Kennett, Li Jin, Hui Li, Johannes Krause, Ron Pinhasi, David Reich

doi: https://doi.org/10.1101/2020.03.25.004606

Abstract

The deep population history of East Asia remains poorly understood due to a lack of ancient DNA data and sparse sampling of present-day people. We report genome-wide data from 191 individuals from Mongolia, northern China, Taiwan, the Amur River Basin and Japan dating to 6000 BCE - 1000 CE, many from contexts never previously analyzed with ancient DNA. We also report 383 present-day individuals from 46 groups mostly from the Tibetan Plateau and southern China. We document how 6000-3600 BCE people of Mongolia and the Amur River Basin were from populations that expanded over Northeast Asia, likely dispersing the ancestors of Mongolic and Tungusic languages. In a time transect of 89 Mongolians, we reveal how Yamnaya steppe pastoralist spread from the west by 3300-2900 BCE in association with the Afanasievo culture, although we also document a boy buried in an Afanasievo barrow with ancestry entirely from local Mongolian hunter-gatherers, representing a unique case of someone of entirely non-Yamnaya ancestry interred in this way. The second spread of Yamnaya-derived ancestry came via groups that harbored about a third of their ancestry from European farmers, which nearly completely displaced unmixed Yamnaya-related lineages in Mongolia in the second millennium BCE, but did not replace Afanasievo lineages in western China where Afanasievo ancestry persisted, plausibly acting as the source of the early-splitting Tocharian branch of Indo-European languages. Analyzing 20 Yellow River Basin farmers dating to ~3000 BCE, we document a population that was a plausible vector for the spread of Sino-Tibetan languages both to the Tibetan Plateau and to the central plain where they mixed with southern agriculturalists to form the ancestors of Han Chinese. We show that the individuals in a time transect of 52 ancient Taiwan individuals spanning at least 1400 BCE to 600 CE were consistent with being nearly direct descendants of Yangtze Valley first farmers who likely spread Austronesian, Tai-Kadai and Austroasiatic languages across Southeast and South Asia and mixing with the people they encountered, contributing to a four-fold reduction of genetic differentiation during the emergence of complex societies. We finally report data from Jomon hunter-gatherers from Japan who harbored one of the earliest splitting branches of East Eurasian variation, and show an affinity among Jomon, Amur River Basin, ancient Taiwan, and Austronesian-speakers, as expected for ancestry if they all had contributions from a Late Pleistocene coastal route migration to East Asia.

Глубокая популяционная история Восточной Азии остается малоизученной из-за отсутствия данных древней ДНК и скудной выборки современных людей. Мы сообщаем данные по всему геному от 191 человека из Монголии, северного Китая, Тайваня, бассейна реки Амур и Японии, датируемые 6000 г. до н.э. - 1000 г. н.э., многие из контекстов, которые ранее никогда не анализировались с помощью древней ДНК. Мы также сообщаем о 383 современных индивидуумах из 46 групп, в основном с Тибетского нагорья и южного Китая. Мы документируем, как в период 6000-3600 до н.э. жители Монголии и бассейна реки Амур были из населения, которое распространилось по Северо-Восточной Азии, вероятно, распространяя предков монгольских и тунгусских народов. Во временном разрезе 89 представителей древнего населения Монголии мы мы показываем, как к 3300-2900 гг. до н.э. ямные степные скотоводы распространились с запада в связи с Афанасьевской культурой, хотя мы также документируем мальчика, похороненного в Афанасьевском кургане с происхождением исключительно от местных монгольских охотников-собирателей , представляющий уникальный случай, когда кто-то совершенно не-ямного происхождения был похоронен в соответствии с ямным обрядом. Второе распространение ямного генетического компонента произошло через группы, которые унаследовали около трети своих генов от европейских земледельцев, и которые во втором тысячелетии до н.э. почти полностью вытеснили несмешанных потомков ямников в Монголии, но не вытеснили потомков афанасьевцев в западном Китае, где афанасьевский генетический компонент сохранился, возможно  выступив источником рано отделившейся тохарской ветви индоевропейских языков. Анализируя 20 земледельцев бассейна реки Хуанхэ, датируемых примерно 3000 годом до н. э., мы фиксируем население, которое было вероятным вектором распространения сино-тибетских языков как на Тибетское плато, так и на центральную равнину, где они смешались с южными земледельцами, чтобы сформировать предков ханьцев(китайцев). Мы показываем, что индивидуумы во временном разрезе 52 древних тайваньцев, охватывающем, по крайней мере, 1400 г. до н.э. - 600 г. н.э., соответствовали почти прямым потомкам первых земледельцев долины Янцзы, которые, вероятно, распространили австронезийские, Тай-Кадайские и Австроазиатские языки по Юго-Восточной и Южной Азии и смешавшись с людьми, с которыми они столкнулись, поспособствовали четырехкратному снижению генетической дифференциации в период возникновения сложных обществ. Наконец, мы приводим данные дзёмонских охотников-собирателей из Японии, которые представляют одну из самых ранних отделившихся ветвей восточно-евразийской вариации, и показываем сходство между населением дзёмона, бассейном реки Амур, древним Тайванем и Австронезийцами, как и ожидалось в плане генетики, если бы все они имели вклад от позднеплейстоценовой миграции по прибрежному пути в Восточную Азию.

Online Table 1: Newly reported ancient individuals

Master ID Date: One of two formats. (Format 1) 95.4% CI calibrated radiocarbon age (Conventional Radiocarbon Age BP, Lab number) e.g. 5983-5747 calBCE (6980±50 BP, Beta-226472). (Format 2) Archaeological context date, e.g. 2500-1700 BCE Locality Region Y chromosome haplogroup (called only for males with 1240K data and only if >50000 autosomal SNPs hit) mtDNA haplogroup only called if >2x coverage
AH1-17 2904-2862 calBCE (4230±25 BP, XA-15378) Wuzhuangguoliang, Shaanxi, China China n/a (female) ..
18R21263 3400-2800 BCE Wuzhuangguoliang, Shaanxi, China China n/a (exome capture) n/a (exome capture)
S91 3400-2800 BCE Wuzhuangguoliang, Shaanxi, China China n/a (female) n/a (exome capture)
S118 3400-2800 BCE Wuzhuangguoliang, Shaanxi, China China n/a (female) n/a (exome capture)
S94 3400-2800 BCE Wuzhuangguoliang, Shaanxi, China China n/a (female) n/a (exome capture)
18R21262 3400-2800 BCE Wuzhuangguoliang, Shaanxi, China China n/a (female) n/a (exome capture)
18R21264 3400-2800 BCE Wuzhuangguoliang, Shaanxi, China China n/a (female) n/a (exome capture)
S123 3400-2800 BCE Wuzhuangguoliang, Shaanxi, China China n/a (female) n/a (exome capture)
18R21266 3400-2800 BCE Wuzhuangguoliang, Shaanxi, China China n/a (female) n/a (exome capture)
18R21268 3400-2800 BCE Wuzhuangguoliang, Shaanxi, China China n/a (female) n/a (exome capture)
S96 3400-2800 BCE Wuzhuangguoliang, Shaanxi, China China n/a (female) n/a (exome capture)
S122 3400-2800 BCE Wuzhuangguoliang, Shaanxi, China China n/a (female) n/a (exome capture)
S120 3400-2800 BCE Wuzhuangguoliang, Shaanxi, China China n/a (exome capture) n/a (exome capture)
S97 3400-2800 BCE Wuzhuangguoliang, Shaanxi, China China n/a (exome capture) n/a (exome capture)
18R21265 3400-2800 BCE Wuzhuangguoliang, Shaanxi, China China n/a (exome capture) n/a (exome capture)
18R21267 3400-2800 BCE Wuzhuangguoliang, Shaanxi, China China n/a (exome capture) n/a (exome capture)
S95 3400-2800 BCE Wuzhuangguoliang, Shaanxi, China China n/a (exome capture) n/a (exome capture)
AH1-7 3348-3090 calBCE (4490±30 BP, XA-15382) Wuzhuangguoliang, Shaanxi, China China n/a (female) ..
S121 3400-2800 BCE Wuzhuangguoliang, Shaanxi, China China n/a (exome capture) n/a (exome capture)
S119 3400-2800 BCE Wuzhuangguoliang, Shaanxi, China China n/a (exome capture) n/a (exome capture)
I13882 2000-1500 BCE Rokutsu Shell Mound Japan n/a (female) N9b
I13883 2000-1500 BCE Rokutsu Shell Mound Japan D1b1d1a N9b2a
I13884 2000-1500 BCE Rokutsu Shell Mound Japan n/a (female) N9b
I13885 2000-1500 BCE Rokutsu Shell Mound Japan n/a (female) N9b
I13886 2000-1500 BCE Rokutsu Shell Mound Japan D1b1d1 N9b1
I13887 2000-1500 BCE Rokutsu Shell Mound Japan D1b1d1 N9b1
I6341 3500-3200 BCE Funadomari Japan D N9b1
I12973 1388-1134 calBCE (3020±25 BP, UCIAMS-226529, PSUG-5432) Bulgan, Khutag undur sum,  Marzyn Khutul tomb 1 Mongolia Q1a2a1c D4j
I12958 1619-1458 calBCE (3265±30 BP, UCIAMS-226527, PSUG-5430) [use instead of less precise and inconsistent non-AMS-date of 1187-516 calBCE (2685±110 BP, Le-7213)] Khovd, Bulgan sum, Khukh uzuur, barrow 2 Mongolia n/a (female) G2a1
I6221 3000-2500 BCE Bayankhongor aimag, Erdenetsogt sum, Shatar chuluu kurgan 2 Mongolia J1 U5a1+@16192
I6222 3316-2918 calBCE (4415±31 BP, OxA-36222) Bayankhongor aimag, Erdenetsogt sum, Shatar chuluu kurgan 2 Mongolia R1b1a1a2a1a R
I13957 2858-2505 calBCE (4095±25 BP, PSUAMS-6942) [use instead of less precise but consistent non-AMS-date of 3011-2480 calBCE (4180±100 BP, Le-7219)] Bayan-Ulgiii aimag, Ulaankhus sum, Kurgak govi barrow 1, infant burial in main grave Mongolia C2b1a1 G2a+152
I13180 3013-2876 calBCE (4290±35 BP, PSUAMS-6944) [we used this direct AMS date on the same petrous bone analyzed for DNA instead of a previous and marginally non-overalpping AMS date of 2866-2577 calBCE (4114±29 BP, OxA-36230) on a different skeletal element supposed to be from the same individual, or a non-AMS date of 2572-2151 calBCE (3900±70 BP, Le-7212) also a different skeletal element supposed to be from the same individual] Bayan-Ulgii aimag, Ulaankhus sum, Kumdii govi barrow, burial 2 Mongolia n/a (female) A+152+16362
I12977 2913-2777 calBCE (4250±25 BP, UCIAMS-226551, PSUG-5466) [use instead of less precise and inconsistent non-AMS date of 2471-1781 calBCE (3725±115 BP, Le-7220)] Bayan-Ulgiii aimag, Ulaankhus sum, Kulala Ula barrow 1, secondary burial Mongolia C2b1a1 D4b1
I6361 2618-2487 calBCE (4034±16 BP, GrM-12938) [use instead of less precise and inconsistent non-AMS date of 2471-2044 calBCE (3825±70 BP, Le-7215)] Bayan-Ulgii aimag, Ulaankhus sum, Kurgak govi barrow 2 secondary burial Mongolia n/a (female) A+152+16362
I12978 2570-2476 calBCE [Rcombine of 2567-2468 calBCE (3983±17 BP, GrM-12984) and 2833-2487 calBCE (4050±25 BP, UCIAMS-226530, PSUG-5433); use instead of less precise and inconsistent non-AMS dates of 2461-2152 calBCE (3838±40 BP, Le-6932, Le-6933, Le-6939 R_Combine)] Khovd, Bulgan sum, Yagshiin hodoo barrow 1, bone find B Mongolia R1b1a1a2a2 C4+152
I12957 2571-2464 calBCE (3980±25 BP, UCIAMS-226526, PSUG-5429) Khovd, Bulgan sum, Yagshiin khodoo, Barrow 3 Mongolia R1b1a1a2a2 T1a1
I6369 752-416 calBCE (2455±20 BP, PSUAMS-3114) Khentii, Murun sum, Bus khairkhan Tomb 2 Mongolia n/a (female) B5b
I6353 1002-909 calBCE [Rcombine of 1002-846 calBCE (2780±25 BP, PSUAMS-2946), 1072-903 calBCE (2826±30 BP, OxA-36456)] Uvurkhangai, Khujirt sum, Shunkhlai Uul, Tomb 9 Mongolia Q1a1a1 G2b2
I12969 1119-980 calBCE (2875±20 BP, UCIAMS-226545, PSUG-5460) Khentii, Delgerkhaan sum, Khanan 3 Mongolia Q1a1a1 F1b1+@152
I13178 1042-916 calBCE (2825±20 BP, UCIAMS-226531, PSUG-5434) Khentii, Bayan-Adraga sum, Shuusyn boom Tomb 6 Mongolia n/a (female) R
I13958 1023-912 calBCE (2820±20 BP, UCIAMS-226558, PSUG-5473) Tuv aimag, Altanbulag sum, Erdenekhairkhan uulyn ar, Tomb 3 Mongolia n/a (<50000 SNPs) H2a
I7032 976-838 calBCE (2765±20 BP, PSUAMS-4004) Khentii, Batshireet sum, Tugsiin uvdug Tomb 1 Mongolia n/a (female) C4a2a1
I12971 976-838 calBCE (2765±20 BP, UCIAMS-226547, PSUG-5462) Dornod aimag, Gurvanzagal sum, Khadan khuv Tomb 1 Mongolia n/a (female) D4j7
I13963 968-833 calBCE (2750±20 BP, PSUAMS-6943) Khentii, Delgerkhaan sum, Rashaant tolgoi, tomb 6 Mongolia n/a (female) C5b1b
I6368 791-569 calBCE [Rcombine of 784-544 calBCE (2510±20 BP, PSUAMS-3907) and 800-571 calBCE (2550±20 BP, PSUAMS-4252)] Khentii, Delgerkhaan sum, Khanan Tomb 1a Mongolia n/a (female) L3
I6349 895-801 calBCE (2675±20 BP, PSUAMS-7398) Sükhbaatar aimag, Dariganga sum, Baaran-Khavtsal Mongolia Q1a1a1 D4m2
I6352 1071-925 calBCE (2845±20 BP, PSUAMS-7276) Uvurkhangai, Khujirt sum, Shunkhlai Uul, Grave #1 Mongolia Q1a1a F1b1b
I6365 814-783 calBCE (2610±20 BP, PSUAMS-7282) Khuvsgul aimag, Burentogtoh sum, Uushigiin uvur, slab grave 1 Mongolia N1c1a D4o1
I6359 1100-400 BCE [range based on other dates from the same cultural context and genetic cluster] Khuvsgul aimag, Alag-Erdene sum, Erhel nuur, Ulaan tolgoi Mongolia Q1a2a1c C5c
I6357 1100-400 BCE [This range is based on other dates from the same cultural context and genetic cluster. We ignore the direct date of 1277-1383 calCE (680±15 BP, PSUAMS-7278) on a post-cranial element for this individual because it mismatches the archaeological chronology as well as the genetic data (which match each other). The post-cranial was sampled from a box separate from the petrous bone analyzed for ancient DNA, which in the context of the mismatch with the genetic and archaeological context makes it plausible that the post-cranial element is not from the individual analyzed for DNA.] Tuv aimag, Altabulag sum, Tariat Brigad Tomb 5 Mongolia n/a (female) D4b2
I6364 1023-912 calBCE (2820±20 BP, PSUAMS-7281) Khovd, Bulgan sum, Uliastain dood denj I, kurgan 4 Mongolia n/a (female) M9a1a1c1
I13504 487-391 calBCE (2360±20 BP, UCIAMS-226556, PSUG-5471) Khuvsgul aimag, Ulaan-Uul sum Mongolia Q1a2a W
I6230 349-97 calBCE (2140±20 BP, PSUAMS-7396) Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery Grave #47, Burial 2 Mongolia Q1a2a1c1 U2e1h
I7029 354-114 calBCE (2155±20 BP, PSUAMS-3877) Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery Grave 53, burial 2 Mongolia Q1a2a1c1 U4a1
I7026 356-177 calBCE (2180±15 BP, PSUAMS-4503) Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery Grave #41, Burial 6 Mongolia n/a (<50000 SNPs) C4a1a+195
I6231 358-174 calBCE (2180±20 BP, PSUAMS-7397) Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery Grave #47, Burial 3 Mongolia Q1a2a1c1 F1b1b
I6226 360-184 calBCE (2190±25 BP, PSUAMS-7270) Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery Grave #35, Burial 2 Mongolia n/a (female) H91
I6224 361-203 calBCE (2210±20 BP, PSUAMS-7268) Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery Grave #33, Burial 3 Mongolia R1a1a1b T2b34
I6233 361-203 calBCE (2210±20 BP, PSUAMS-7272) Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery, Grave #49, Burial 1 Mongolia R1a1a1b2 F1b1+@152
I7023 365-204 calBCE (2215±20 BP, PSUAMS-4001) Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery Grave #31, Burial 6 Mongolia n/a (female) H
I7028 381-208 calBCE (2235±20 BP, PSUAMS-4597) Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery Grave 46, burial 1 Mongolia n/a (female) H2a2a
I6232 387-209 calBCE (2245±20 BP, PSUAMS-7271) Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery Grave #48, Burial 6 Mongolia Q1a2a1c1 U4b1a1a1
I7030 391-209 calBCE (2250±20 BP, PSUAMS-3722) Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery Grave 56 Mongolia R1a1a1b D4b1a2a1
I7022 391-209 calBCE (2250±20 BP, PSUAMS-4000) Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery Grave #11, Burial 7 Mongolia Q1a1b1 C4a1a+195
I7024 391-209 calBCE (2250±20 BP, PSUAMS-4002) Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery Grave #36, Burial 2 Mongolia R1a1a1b2 R1b1
I7027 399-235 calBCE (2275±20 BP, PSUAMS-4003) [used the PSUAMS date on the same skeletal element analyzed for ancient DNA instead of the non-overlapping measurement on an independent sample from the same individual of 358-195 calBCE (2188±15 BP, GrM-12941) published in Wilkins et al. Nature Ecology and Evolution 2020] Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery Grave #45, Burial 2 Mongolia R1a1a1b T1a1
I6223 400-200 BCE Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery Grave 1 burial 5 Mongolia n/a (<50000 SNPs) ..
I6225 400-200 BCE [This range is based on other dates from the same cultural context and a brother from the same genetic cluster. We ignore the direct date of 1299-1404 calCE (605±20 BP, PSUAMS-7269) on a post-cranial element for this individual because it mismatches the archaeological chronology as well as the genetic data (which match each other). The post-cranial was sampled from a box separate from the petrous bone analyzed for ancient DNA, which in the context of the mismatch with the genetic and archaeological context makes it plausible that the post-cranial element is not from the individual analyzed for DNA.] Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery Grave #33, Burial 5 Mongolia R1a1a T2b34
I6356 400-200 BCE [This range is based on other dates from the same cultural context and genetic cluster. We ignore the direct date of 1386-1217 calBCE (3030±20 BP, PSUAMS-7277) on a mandible for this individual because it mismatches the archaeological chronology as well as the genetic data (which match each other). The mandible sampled for DNA  was different from the petrous bone used for genetic analysis; these skeletal elements were from a commingled burial so it is plausible that the date does not correspond to the genetically analyzed individual.] Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery, Grave #58, Burial 2 Mongolia Q1a2a C4a1a3
I12970 400-235 calBCE (2280±20 BP, UCIAMS-226546, PSUG-5461) Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery Grave 48 burial 7 Mongolia Q1a2a1c1 A8a
I12956 405-364 calBCE (2305±20 BP, UCIAMS-226544, PSUG-5459) Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery Grave 48 burial 4 Mongolia n/a (<50000 SNPs) n/a (<2x)
I13965 397-233 calBCE (2270±20 BP, UCIAMS-226560, PSUG-5475) Bayan-Ulgiii aimag, Tsengel sum, Biluut 2 barrow 4 Mongolia n/a (female) G2a1
I6263 358-181 calBCE (2185±20 BP, PSUAMS-7273) Bayan Ulgii aimag, Tsengel sum, Tsagaan-Uul, Khuvsgol Mongolia n/a (female) H5a1
I6228 3-120 calCE (1950±20 BP, PSUAMS-7395) Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery Grave #44 Mongolia C2b1a1b1 D4j12
I14194 971-834 calBCE (2755±20 BP, UCIAMS-226562, PSUG-5476) Sükhbaatar aimag, Tuvshinshiree sum, Ulaanzukh,  Tomb 32 Mongolia n/a (female) C4a1
I13175 383-535 calCE (1625±20 BP, UCIAMS-226554, PSUG-5469) Khovd, Bulgan sum, Uliastai deed denj II kurgan 1 burial 1 (secondary) Mongolia R1b1c D4b1a2a
I13174 1500-1423 calBCE (3185±20 BP, UCIAMS-226564, PSUG-5483) [use instead of less precise but consistent non-AMS date of 1493-1091 calBCE (3050±70 BP, Le-6638)] Khovd, Munkhkhairkhan sum, Khukh tolgoi barrow 9 Mongolia Q1a2a1c U5a2
I13959 1441-1306 calBCE (3125±20 BP, UCIAMS-226559, PSUG-5474) [use instead of less precise but consistent non-AMS date of 1972-1306 calBCE (3340±130 BP, Le-6637)] Khovd, Munkhkhairkhan sum, Ulaan tolgoi, barrow 3 Mongolia n/a (female) n/a (<2x)
I12972 1447-1309 calBCE (3130±20 BP, UCIAMS-226553, PSUG-5468) Sükhbaatar aimag, Tuvshinshiree sum, Adgiin gol Tomb 21 Mongolia Q1a1 D4i
I14037 1420-1296 calBCE (3095±20 BP, UCIAMS-226561, PSUG-5475) Sükhbaatar aimag, Tuvshinshiree sum, Bulgiin ekh, Tomb 47 Mongolia Q1a1a M11b
I13964 1500-800 BCE Sükhbaatar aimag, Tuvshinshiree sum, Ulaanzukh, Tomb 17 Mongolia n/a (female) C4a1a+195
I12960 1437-1304 calBCE (3120±20 BP, UCIAMS-226528, PSUG-5431) Uvurkhangai, Khujirt sum, Shunkhlai Uul, Tomb 2 Mongolia Q1a1 C4a1a+195
I12506 1438-1299 calBCE (3115±25 BP, PSUAMS-6933) [use instead of previous date of 1608-1416 calBCE (3215±40 BP, OxA-36459) not from the same skeletal element] Sükhbaatar aimag, Tuvshinshiree sum, Ulaanzukh, Tomb 42 Mongolia n/a (<50000 SNPs) n/a (<2x)
I12976 1437-1304 calBCE (3120±20 BP, UCIAMS-226550, PSUG-5465) [use instead of less precise but consistent non-AMS date of 1396-845 calBCE (2910±110 BP, Le-7226)] Bayan-Ulgiii aimag, Ulaankhus sum, Kulala Ula barrow 2 Mongolia N1 U2e2a1a2
I7033 1209-1038 calBCE (2920±20 BP, PSUAMS-4005) Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery kurgan 59 Mongolia R1a1a1b R
I6363 1214-1058 calBCE [Rcombine of 1210-1047 calBCE (2925±20 BP, PSUAMS-7280) and 1277-1057 calBCE (2967±31 BP, OxA-X-2737-54); do not use the less precise but consistent non-AMS date of 1208-841 calBCE (2840±70 BP, Le-7979)] Khovd, Bulgan sum, Khudzrtyn gol II, barrow 1, burial 4 (main burial) Mongolia R1a1a1b2a2b U2e1
I6351 1003-854 calBCE (2785±20 BP, PSUAMS-7275) Uvs aimag, Khyargas sum, Urgun shireg, Tomb 3 Mongolia n/a (female) C4b1
I13505 1119-980 calBCE (2875±20 BP, UCIAMS-226557, PSUG-5472) Bayankhongor aimag, Erdentsogt sum Mongolia Q1a1a1 A+152+16362
I7039 1209-1038 calBCE (2920±20 BP, PSUAMS-4006) Khovd, Duut sum, Khoit Tsenkheriin gol 1988 barrow 3 Mongolia Q1a2a1c D4j
I13768 1210-1047 calBCE (2925±20 BP, PSUAMS-7115) Bayankhongor aimag, Ulziit sum, Muunit uul Mongolia N D4j1
I12975 1226-1056 calBCE (2950±20 BP, UCIAMS-226549, PSUG-5464) Khuvsgul aimag, Khankh sum, Khavtsal II Tomb 1 (Khirgisuur) Mongolia C2b1a U5a1a2a
I13767 1374-1131 calBCE (3005±20 BP, UCIAMS-226534, PSUG-5437) Bayankhongor aimag, Ulziit sum, Khar Sair am Mongolia Q1a2a1c R1b1
I13766 1381-1131 calBCE (3010±25 BP, UCIAMS-226533, PSUG-5436) Bayankhongor aimag, Erdentsogt sum Mongolia Q1a2a1c C5b1b
I6262 1396-1231 calBCE (3050±20 BP, PSUAMS-3111) Bayankhongor aimag, Khar Sairiin Am, Khirigsuur Mongolia n/a (female) A8a1
I6264 1397-1236 calBCE (3055±20 BP, PSUAMS-3112) Bayankhongor aimag, Khar Sairiin Am, Khirigsuur Mongolia n/a (female) A8a1
I6347 1411-1284 calBCE (3080±20 BP, PSUAMS-3113) Uvs aimag, Züünkhangai sum, Savaa Khar Mongolia n/a (female) K2a5
I6362 1015-911 calBCE (2815±20 BP, PSUAMS-2955) Khovd, Bulgan sum, Khudzhirtyn gol I, kurgan 2 Mongolia n/a (female) D4b1
I6367 1258-1088 calBCE (2955±20 BP, PSUAMS-7284) [use instead of less precise but agreeing date of 1260-1040 calBCE (2820±30 BP, Beta-306037)] Bayan-Ulgii aimag, Tsengel sum, Biluut 2, barrow 3 Mongolia O2a2b1a1a C4a1
I12955 1948-1777 calBCE (3545±20 BP, UCIAMS-226543, PSUG-5458) Zavkhan, Bayantes sum, Khukh Khushoony bom, Barrow 3 Mongolia N1c1a D4i
I6348 1746-1630 calBCE (3395±25 BP, PSUAMS-7274) Bulgan, Khutag-Undur sum, Haruulyn Gozgor tomb 1-113 Mongolia Q1a2a1c HV12b1
I13173 1879-1694 calBCE (3460±20 BP, UCIAMS-226552, PSUG-5467) [use instead of less precise and inconsistent non-AMS date of 1609-1232 calBCE (3150±70 BP, Le-6936)] Khovd, Munkhkhairkhan sum, Ulan goviin uzuur barrow 2 Mongolia N H15b1
I13176 890-982 calCE (1115±20 BP, UCIAMS-226555, PSUG-5470) Khovd, Bulgan sum, Hudzhirtyn gol II, kurgan 1 burial 3 (secondary) Mongolia R1b1c D4i
I12974 1225-1285 calCE (750±20 BP, UCIAMS-226548, PSUG-5463) Khentii, Murun sum, Bus khairkhan Tomb 3 Mongolia n/a (female) C4a1a+195
I13961 1282-1388 calCE (660±15 BP, PSUAMS-7114) Dornod aimag, Gurvanzagal sum  Tsagaan chuluut kurgan 1 Mongolia n/a (female) A+152+16362
I7021 5211-5000 calBCE (6145±35 BP, PSUAMS-3931) Dornod aimag, Bulgan sum, Norovlin Uul burial Mongolia C2b1a1 D4h4
I7031 4458-4364 calBCE (5590±20 BP, PSUAMS-4598) Dornod aimag, Bayantumen sum, Kherlengiin baruun ereg (Kerulen River right bank), Grave 2 (1968 y.) Mongolia n/a (<50000 SNPs) C5b1
I6358 5974-5747 calBCE (6965±35 BP, PSUAMS-7279) Dornod aimag, Bayantumen sum, Kherlen River South bank nearby Choibalsan town Mongolia n/a (female) F1b1+@152
I13698 5700-5000 BCE Bulgan, Khutag-Undur sum, Egiin-gol river valley, Marzyn Khutul, tomb 1-055-1, sceleton 2 Mongolia C2b1a1 F1b1+@152
I13701 5721-5614 calBCE (6739±40 BP, AA-107617) Bulgan, Khutag-Undur sum, Egiin-gol river valley, Kharuulyn gozgor, tomb 1-116 Mongolia n/a (female) C4a1a3
I13179 5607-5477 calBCE (6565±30 BP, UCIAMS-226532, PSUG-5435) Bulgan, Khutag undur sum, Kharuulyn gozgor, tomb 1-116 Mongolia n/a (female) C4a1a3
I14000 5575-5474 calBCE (6571±47 BP, COL3897.1.1) Bulgan, Khutag-Undur sum, Egiin-gol river valley, Kharuulyn gozgor, tomb 1-117-1, sceleton 3 Mongolia n/a (female) D4j
I11696 5632-5531 calBCE (6655±30 BP, UCIAMS-226523, PSUG-5426) Bulgan, Khutag-Undur sum,  Marzyn khutul, Tomb 1-055-1, sceleton 1 Mongolia C2b1a1 D4j
I11698 5632-5515 calBCE (6645±35 BP, UCIAMS-226525, PSUG-5428) Bulgan, Khutag-Undur sum,  Marzyn khutul, Tomb 1-055-1, sceleton 2 Mongolia C2b1a1 D4b1c
I11697 5617-5491 calBCE (6610±30 BP, UCIAMS-226524, PSUG-5427) Bulgan, Khutag-Undur sum,  Marzyn khutul, Tomb 1-055-1, sceleton 4 Mongolia C2b1a1 D4j8
I6366 1410-1261 calBCE (3065±25 BP, PSUAMS-7283) Bayan-Ulgiii aimag, Tsengel sum, Biluut 2 barrow 2 Mongolia n/a (female) J2b1a2a
I1208 450-750 CE Primorsky Krai, Chernyatino 5, Mohe-Bohai Graveyard Russia n/a (<50000 SNPs) H2
I14819 3750-3600 BCE [layer date based on mothering children with I3356 at 3696-3638 calBCE (4875±20 BP, PSUAMS-2160)] Primorsky Krai, Boisman-2 Russia n/a (female) C5a
I14308 5400-3600 BCE Primorsky Krai, Boisman-2 Russia C2a1a2(xC2a1a2a) D4o2a
I3356 3696-3638 calBCE (4875±20 BP, PSUAMS-2160) Primorsky Krai, Boisman-2 Russia C2a1a3(xC2a1a3a) C5b1
I3354 5371-5219 calBCE (6325±35 BP, PSUAMS-2070) Primorsky Krai, Boisman-2 Russia n/a (female) D4
I1196 5213-5008 calBCE (6155±35 BP, Poz-83476) Primorsky Krai, Boisman-2 Russia n/a (female) C5b1
I1206 5213-4834 calBCE (6080±70 BP, TERRA-b06129705) Primorsky Krai, Boisman-2 Russia n/a (female) R
I1198 5034-4806 calBCE (6030±40 BP, Poz-83478) Primorsky Krai, Boisman-2 Russia n/a (female) D4l2
I1193 4981-4795 calBCE (5995±35 BP, Poz-83474) Primorsky Krai, Boisman-2 Russia C2a1a3 C5b
I3355 4981-4787 calBCE (5985±35 BP, PSUAMS-2114) Primorsky Krai, Boisman-2 Russia C2a1a(xC2a1a1,C2a1a2,C2a1a3) C5a
I1197 4956-4729 calBCE (5970±40 BP, Poz-83477) Primorsky Krai, Boisman-2 Russia n/a (female) C5a
I1194 4792-4604 calBCE (5840±35 BP, Poz-83475) Primorsky Krai, Boisman-2 Russia n/a (female) C5a
I1190 4721-4335 calBCE (5860±100 BP, TERRA-06129717) Primorsky Krai, Boisman-2 Russia n/a (female) D4e5
I1192 4932-4608 calBCE (5890±60 BP, TERRA-b06129733) Primorsky Krai, Boisman-2 Russia C2a1(xC2a1a1a,C2a1a2a) B4d1'2'3
I14307 5400-3600 BCE Primorsky Krai, Boisman-2 Russia n/a (female) D4o2a
I14772 3750-3600 BCE [layer date based on being daughter of I3356 at 3696-3638 calBCE (4875±20 BP, PSUAMS-2160)] Primorsky Krai, Boisman-2 Russia n/a (female) C5a
I14774 3750-3600 BCE [layer date based on being daughter of I3356 at 3696-3638 calBCE (4875±20 BP, PSUAMS-2160)] Primorsky Krai, Boisman-2 Russia n/a (female) C5a
I14773 3750-3600 BCE [layer date based on being son of I3356 at 3696-3638 calBCE (4875±20 BP, PSUAMS-2160)] Primorsky Krai, Boisman-2 Russia n/a (<50000 SNPs) C5a
I14771 3750-3600 BCE [layer date based on being son of I3356 at 3696-3638 calBCE (4875±20 BP, PSUAMS-2160)] Primorsky Krai, Boisman-2 Russia C2a(xC2a1a1,C2a1a2a) C5a
I1209 1039-1224 calCE (880±35 BP, Poz-83439) Roshino-4, Primorsky Krai Russia O2 D4a1e
I3358 1050-1220 calCE (875±20 BP, PSUAMS-2115) Roshino-4, Primorsky Krai Russia n/a (female) D4a1
I15926 1800-1950 CE Nevelsk 2 Russia D1a2b(xD1a2b1) N9b1
I1202 996-830 calBCE (2760±35 BP, Poz-83438) Pospelovo 1, Primorsky Krai Russia N1a(xN1a1,N1a2) C5b
I13721 250-650 CE Green Island, Gongguan site Taiwan n/a (female) Y2a1
I13722 250-650 CE Green Island, Gongguan site Taiwan n/a (<50000 SNPs) Y2a1
I3618 1402-1262 calBCE (3060±20 BP, PSUAMS-2268) Yilan, Hanben Taiwan O2a2b2a2 R
I8071 150-465 CE Yilan, Hanben Taiwan n/a (female) R
I13695 150-465 CE Yilan, Hanben Taiwan n/a (<50000 SNPs) F4b
I8076 150-465 CE Yilan, Hanben Taiwan n/a (<50000 SNPs) R30
I8080 150-465 CE Yilan, Hanben Taiwan O1a1a1a F3b1a+16093
I8075 150-465 CE Yilan, Hanben Taiwan n/a (female) E2b
I3611 150-465 CE Yilan, Hanben Taiwan n/a (female) E1a1a1
I3616 150-465 CE Yilan, Hanben Taiwan n/a (female) B4b1a2
I3615 150-465 CE Yilan, Hanben Taiwan n/a (female) R
I3620 150-465 CE Yilan, Hanben Taiwan n/a (female) R
I3617 150-465 CE Yilan, Hanben Taiwan n/a (female) F4b1
I3621 150-465 CE Yilan, Hanben Taiwan n/a (female) R9c1b2
I15159 150-465 CE Yilan, Hanben Taiwan n/a (female) F4b1
I3614 150-465 CE Yilan, Hanben Taiwan O2a2b2a R
I3612 150-465 CE Yilan, Hanben Taiwan O2a2b2a2 E1a1a1
I3613 150-465 CE Yilan, Hanben Taiwan O1a1a1a1 R
I3619 395-536 calCE (1610±25 BP, PSUAMS-2075) Yilan, Hanben Taiwan n/a (female) F3b1a+16093
I14925 510-870 CE Yilan, Hanben Taiwan n/a (female) M7b1a2a1
I14934 510-870 CE Yilan, Hanben Taiwan O1a1a1a1 E2a
I14933 510-870 CE Yilan, Hanben Taiwan O1a2 F3b1a
I14931 510-870 CE Yilan, Hanben Taiwan O2a2 E1a1a1
I8072 510-870 CE Yilan, HanBen Taiwan O1a E1a1a
I14929 510-870 CE Yilan, Hanben Taiwan O2a2b B4a1a
I8074 510-870 CE Yilan, Hanben Taiwan n/a (female) F3b1a+16093
I8081 510-870 CE Yilan, Hanben Taiwan O1a1a1a F3b1a2
I15158 510-870 CE Yilan, Hanben Taiwan O1a2 B4b1a2f
I13692 510-870 CE Yilan, Hanben Taiwan n/a (female) M7c1c3
I3728 510-870 CE Yilan, Hanben Taiwan n/a (female) F3b1a+16093
I3736 510-870 CE Yilan, Hanben Taiwan O1a1a1a1 F4b1
I3732 510-870 CE Yilan, Hanben Taiwan n/a (female) F1a3a
I3735 510-870 CE Yilan, Hanben Taiwan n/a (female) E1a
I3727 510-870 CE Yilan, Hanben Taiwan n/a (female) D6a2
I14937 510-870 CE Yilan, Hanben Taiwan n/a (female) B4a1a
I3734 510-870 CE Yilan, Hanben Taiwan O2a2a1a2 E1a
I3731 510-870 CE Yilan, Hanben Taiwan O1a1a1a1a1 F3b1
I3733 510-870 CE Yilan, Hanben Taiwan O1a1a1a1 F3b1
I8078 510-870 CE Yilan, Hanben Taiwan O2a2b2 E1a1
I15156 510-870 CE Yilan, Hanben Taiwan n/a (<50000 SNPs) D6a2
I13697 150-465 CE Yilan, Hanben Taiwan n/a (<50000 SNPs) R
I15164 150-465 CE Yilan, Hanben Taiwan n/a (female) E2a
I15160 150-465 CE Yilan, Hanben Taiwan n/a (<50000 SNPs) F4b1
I15162 150-465 CE Yilan, Hanben Taiwan n/a (<50000 SNPs) E1a1a
I15166 150-465 CE Yilan, Hanben Taiwan n/a (female) E1a1a
I8079 150-465 CE Yilan, Hanben Taiwan n/a (female) F3b1
I14930 510-870 CE Yilan, Hanben Taiwan n/a (female) E1a1a1
I14928 510-870 CE Yilan, Hanben Taiwan n/a (female) B5a2a1+16129
I14926 510-870 CE Yilan, Hanben Taiwan n/a (female) F4b1
I13694 510-870 CE Yilan, Hanben Taiwan n/a (female) B4a1a
I14927 510-870 CE Yilan, Hanben Taiwan n/a (female) M7b1a2a1
I13696 510-870 CE Yilan, Hanben Taiwan n/a (<50000 SNPs) n/a (<2x)

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I6222   AT_26, Grave #2, MONU_26, Grave #2   Pinhasi, Ron; Cheronet, Olivia; Sirak, Kendra;  Dashtseveg, Tumen   petrous (CBD)   5067   3316-2918 calBCE (4415±31 BP, OxA-36222)   Confident - The genetic analysis was done on a different individuals from the sample used to produce the direct 14C date (published in Wilkins et al. Nature Ecology and Evolution 2020). However, the 14C date matches to the Afanasievo period, and this individual clusters genetically with previously reported individuals from the Afansievo culture from the Altai region of Russia, providing confidence int he cultural assignment.   Mongolia_Chalcolithic_1_Afanasievo   Yes   Bayankhongor aimag, Erdenetsogt sum, Shatar chuluu kurgan 2   Mongolia   46,4   100,82   1240K capture   1   0,136   146406   M   R1b1a1a2a1a    25    R

По указанной в статье версии ISOGG R1b1a1a2a1a - это L151.

I12978   AT_590B, 707   Pinhasi, Ron; Cheronet, Olivia; Sirak, Kendra;  Tumen, Dashtseveg; Erdenebaatar, Diimaajav; Kovalev, Alexey   petrous   4473   2570-2476 calBCE [Rcombine of 2567-2468 calBCE (3983±17 BP, GrM-12984) and 2833-2487 calBCE (4050±25 BP, UCIAMS-226530, PSUG-5433); use instead of less precise and inconsistent non-AMS dates of 2461-2152 calBCE (3838±40 BP, Le-6932, Le-6933, Le-6939 R_Combine)]   Confident - The genetic data and direct 14C date are from the same exported petrous bone; the direct date and archaeological context (Chemurchek EBA culture) match up; and the genetic patterns place this individual into a cluster of 2 individuals from a Chemuchek EBA archaeological context both of whom are directly dated to a consistent period.    Mongolia_EBA_2_Chemurchek   Yes   Khovd, Bulgan sum, Yagshiin hodoo barrow 1, bone find B   Mongolia   46,12   91,57   1240K capture   1   2,471405   796172   M   R1b1a1a2a2    479    C4+152

I12957   AT_598, 682   Pinhasi, Ron; Cheronet, Olivia; Sirak, Kendra;  Tumen, Dashtseveg; Erdenebaatar, Diimaajav; Kovalev, Alexey   petrous   4468   2571-2464 calBCE (3980±25 BP, UCIAMS-226526, PSUG-5429)   Confident - The genetic data and direct 14C date are from the same exported petrous bone; the direct date and archaeological context (Chemurchek EBA culture) match up; and the genetic patterns place this individual into a cluster of 2 individuals from a Chemuchek EBA archaeological context both of whom are directly dated to a consistent period.    Mongolia_EBA_2_Chemurchek   Yes   Khovd, Bulgan sum, Yagshiin khodoo, Barrow 3   Mongolia   46,12   91,57   1240K capture   1   3,836698   841251   M   R1b1a1a2a2    534    T1a1

R1b1a1a2a2 - это Z2103.
Чемурчекская культура

I13175   AT_679, 882   Pinhasi, Ron; Cheronet, Olivia; Sirak, Kendra;  Tumen, Dashtseveg; Erdenebaatar, Diimaajav; Kovalev, Alexey   petrous   1491   383-535 calCE (1625±20 BP, UCIAMS-226554, PSUG-5469)   Confident - The genetic data and direct 14C date are from the same exported petrous bone. While this individual is the only representative of their genetic cluster, the archaeological context is consistent with the direct date.   Mongolia_IA_Xianbei   Yes   Khovd, Bulgan sum, Uliastai deed denj II kurgan 1 burial 1 (secondary)   Mongolia   45,39   90,798   1240K capture   1   5,534433   848539   M   R1b1c    515    D4b1a2a

I13176   AT_678, 883   Pinhasi, Ron; Cheronet, Olivia; Sirak, Kendra;  Tumen, Dashtseveg; Erdenebaatar, Diimaajav; Kovalev, Alexey   petrous   1014   890-982 calCE (1115±20 BP, UCIAMS-226555, PSUG-5470)   SOME DOUBT - The genetic data and direct 14C date are from the same exported petrous bone. However, this individual is the only representative of their genetic cluster and the archaeological context is not fully clear, so we treat this archaeological and chronological context assignment as tentative.   Mongolia_Medieval   Yes   Khovd, Bulgan sum, Hudzhirtyn gol II, kurgan 1 burial 3 (secondary)   Mongolia   45,35   90,85   1240K capture   1   0,79158   546668   M   R1b1c    151    D4i

R1b1c - это PH155.
https://www.yfull.com/tree/R-PH155/
« Последнее редактирование: 26 Март 2020, 06:41:14 от Arthwr »

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Afanasievo:

I6221   AT_25, Grave #5, MONU_25, Grave #5   Pinhasi, Ron; Cheronet, Olivia; Sirak, Kendra;  Dashtseveg, Tumen   petrous (CBD)   4700   3000-2500 BCE   Confident - Although we do not have a direct 14C date, this individual clusters genetically with previously reported individuals from the Afansievo culture from the Altai region of Russia and so the archaeological assignment is supported by an independent line of evidence.   Mongolia_Chalcolithic_1_Afanasievo   Yes   Bayankhongor aimag, Erdenetsogt sum, Shatar chuluu kurgan 2   Mongolia   46,4   100,82   1240K capture   1   4,338   822131   M   J1    420    U5a1+@16192

I6222   AT_26, Grave #2, MONU_26, Grave #2   Pinhasi, Ron; Cheronet, Olivia; Sirak, Kendra;  Dashtseveg, Tumen   petrous (CBD)   5067   3316-2918 calBCE (4415±31 BP, OxA-36222)   Confident - The genetic analysis was done on a different individuals from the sample used to produce the direct 14C date (published in Wilkins et al. Nature Ecology and Evolution 2020). However, the 14C date matches to the Afanasievo period, and this individual clusters genetically with previously reported individuals from the Afansievo culture from the Altai region of Russia, providing confidence int he cultural assignment.   Mongolia_Chalcolithic_1_Afanasievo   Yes   Bayankhongor aimag, Erdenetsogt sum, Shatar chuluu kurgan 2   Mongolia   46,4   100,82   1240K capture   1   0,136   146406   M   R1b1a1a2a1a    25    R

I13957   AT_629, 990   Pinhasi, Ron; Cheronet, Olivia; Sirak, Kendra;  Tumen, Dashtseveg; Erdenebaatar, Diimaajav; Kovalev, Alexey   petrous   4632   2858-2505 calBCE (4095±25 BP, PSUAMS-6942) [use instead of less precise but consistent non-AMS-date of 3011-2480 calBCE (4180±100 BP, Le-7219)]   Confident - The genetic data and direct 14C date are from the same exported left petrous bone, and the photo of the skull in our records matches the photo of the skeleton in the archaeological literature of an individual estimated to be 5-6 years old. The newly generated 14C date generated at the PSUAMS lab on the same sample used for DNA is also consistent with the previously reported 14C date supposed to come from this individual at the Le lab (Institute of the History of Material Culture, Russian Academy of Sciences), further increasing confidence in the chronology.   Mongolia_Chalcolithic_2_Afanasievo   Yes   Bayan-Ulgiii aimag, Ulaankhus sum, Kurgak govi barrow 1, infant burial in main grave    Mongolia   49,336   88,712   1240K capture   1   3,053235   784274   M   C2b1a1    505    G2a+152

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Afanasievo:

I6221   AT_25, Grave #5, MONU_25, Grave #5   Pinhasi, Ron; Cheronet, Olivia; Sirak, Kendra;  Dashtseveg, Tumen   petrous (CBD)   4700   3000-2500 BCE   Confident - Although we do not have a direct 14C date, this individual clusters genetically with previously reported individuals from the Afansievo culture from the Altai region of Russia and so the archaeological assignment is supported by an independent line of evidence.   Mongolia_Chalcolithic_1_Afanasievo   Yes   Bayankhongor aimag, Erdenetsogt sum, Shatar chuluu kurgan 2   Mongolia   46,4   100,82   1240K capture   1   4,338   822131   M   J1    420    U5a1+@16192

У него и мито как у меня.  :)

Известно ли какой субклад J1?

Оффлайн vvw1717

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 Сэмпл I13957 AT_629, 990 этот вероятно от того; мальчика монголоида  Y-C2b1a1 мт. G2a+152?

Оффлайн Fire

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J1 субклад интересен, если J1c или другой не-Кавказский реликт то это может быть потомок
до-Афанасьевских животноводов центральной Азии. Если Кавказская ветвь то наверно от ранних восточных Ямников или поздних южных Хвалынцев

Оффлайн kardes

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А эти Улангомы и Чандманы кто?
 I6230   349-97 calBCE (2140±20 BP, PSUAMS-7396)   Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery Grave #47, Burial 2   Mongolia   Q1a2a1c1   U2e1h
I7029   354-114 calBCE (2155±20 BP, PSUAMS-3877)   Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery Grave 53, burial 2   Mongolia   Q1a2a1c1   U4a1
I7026   356-177 calBCE (2180±15 BP, PSUAMS-4503)   Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery Grave #41, Burial 6   Mongolia   n/a (<50000 SNPs)   C4a1a+195
I6231   358-174 calBCE (2180±20 BP, PSUAMS-7397)   Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery Grave #47, Burial 3   Mongolia   Q1a2a1c1   F1b1b
I6226   360-184 calBCE (2190±25 BP, PSUAMS-7270)   Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery Grave #35, Burial 2   Mongolia   n/a (female)   H91
I6224   361-203 calBCE (2210±20 BP, PSUAMS-7268)   Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery Grave #33, Burial 3   Mongolia   R1a1a1b   T2b34
I6233   361-203 calBCE (2210±20 BP, PSUAMS-7272)   Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery, Grave #49, Burial 1   Mongolia   R1a1a1b2   F1b1+@152
I7023   365-204 calBCE (2215±20 BP, PSUAMS-4001)   Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery Grave #31, Burial 6    Mongolia   n/a (female)   H
I7028   381-208 calBCE (2235±20 BP, PSUAMS-4597)   Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery Grave 46, burial 1   Mongolia   n/a (female)   H2a2a
I6232   387-209 calBCE (2245±20 BP, PSUAMS-7271)   Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery Grave #48, Burial 6   Mongolia   Q1a2a1c1   U4b1a1a1
I7030   391-209 calBCE (2250±20 BP, PSUAMS-3722)   Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery Grave 56    Mongolia   R1a1a1b   D4b1a2a1
I7022   391-209 calBCE (2250±20 BP, PSUAMS-4000)   Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery Grave #11, Burial 7   Mongolia   Q1a1b1   C4a1a+195
I7024   391-209 calBCE (2250±20 BP, PSUAMS-4002)   Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery Grave #36, Burial 2   Mongolia   R1a1a1b2   R1b1
I7027   399-235 calBCE (2275±20 BP, PSUAMS-4003) [used the PSUAMS date on the same skeletal element analyzed for ancient DNA instead of the non-overlapping measurement on an independent sample from the same individual of 358-195 calBCE (2188±15 BP, GrM-12941) published in Wilkins et al. Nature Ecology and Evolution 2020]   Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery Grave #45, Burial 2   Mongolia   R1a1a1b   T1a1
I6223   400-200 BCE   Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery Grave 1 burial 5   Mongolia   n/a (<50000 SNPs)   ..
I6225   400-200 BCE [This range is based on other dates from the same cultural context and a brother from the same genetic cluster. We ignore the direct date of 1299-1404 calCE (605±20 BP, PSUAMS-7269) on a post-cranial element for this individual because it mismatches the archaeological chronology as well as the genetic data (which match each other). The post-cranial was sampled from a box separate from the petrous bone analyzed for ancient DNA, which in the context of the mismatch with the genetic and archaeological context makes it plausible that the post-cranial element is not from the individual analyzed for DNA.]    Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery Grave #33, Burial 5   Mongolia   R1a1a   T2b34
I6356   400-200 BCE [This range is based on other dates from the same cultural context and genetic cluster. We ignore the direct date of 1386-1217 calBCE (3030±20 BP, PSUAMS-7277) on a mandible for this individual because it mismatches the archaeological chronology as well as the genetic data (which match each other). The mandible sampled for DNA  was different from the petrous bone used for genetic analysis; these skeletal elements were from a commingled burial so it is plausible that the date does not correspond to the genetically analyzed individual.]    Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery, Grave #58, Burial 2   Mongolia   Q1a2a   C4a1a3
I12970   400-235 calBCE (2280±20 BP, UCIAMS-226546, PSUG-5461)   Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery Grave 48 burial 7   Mongolia   Q1a2a1c1   A8a
I12956   405-364 calBCE (2305±20 BP, UCIAMS-226544, PSUG-5459)   Uvs aimag, Ulaangom sum, Chandman Uul, Ulaangom cemetery Grave 48 burial 4   Mongolia   n/a (<50000 SNPs)   n/a (<2x)
 

Оффлайн Arame

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  • J1-Z1842
Отличная работа. Наконец что то из восточной Азии.
Связка чемурчекской культуры и афанасьево потверждается. С такими европоидными стеллами они не могли иметь другое происхождение. Вообщем тохарский вопрос практически решен. Ковалев был прав про Чемурчекскую культуру только вот он ошибочно поставил их корни в Центральной Европе. Они имели степное происхождение.
J1 скорее всего из тех реликтовых что были в Карелии в мезолите.
При дальнейшем копании там всплывет также PH155. Который был в Джунгарии в железном веке.

Оффлайн zastrug

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  • I2b1c (P78+)=I2a2a1b1a
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Ковалев был прав про Чемурчекскую культуру только вот он ошибочно поставил их корни в Центральной Европе. Они имели степное происхождение.
Не в Центральной. Он ее истоки видел в неолитическрй культуре мегалитов Южной Франции. Ну то есть там совсем другое, особенно в плане генетики.
Вот кто прав оказался так Напольских - он говорил что в сеймино-турбинском феномене просматриваются следы афанасьевской культуры. Ну а чемурчекская культура точно имела на него влияние, то есть опосредованно афанасьевцы.

Оффлайн Eugene

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По N только Поспелово 1 шт. , и Монголия несколько штук из разных веков.
Но я вижу больше половины не определены гаплы.

Оффлайн User

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I6221   

J1

U5a1+@16192

Если я правильно понимаю, U5a1+@16192 это U5a1i, – редкий субклад данной гаплогруппы был найден в Хвалынской культуре. Поэтому есть вероятность, что J1 окажется J-Z1842.  Ранее в Хвалынской культуре уже была найдена Y-гаплогруппа J.

Оффлайн Arthwr

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I6221   

J1

U5a1+@16192

Если я правильно понимаю, U5a1+@16192 это U5a1i, – редкий субклад данной гаплогруппы был найден в Хвалынской культуре. Поэтому есть вероятность, что J1 окажется J-Z1842.  Ранее в Хвалынской культуре уже была найдена Y-гаплогруппа J.

Вряд ли. Скорее что-то типа:
J1 скорее всего из тех реликтовых что были в Карелии в мезолите.
Так как у образца хорошее покрытие, а нижележащие субклады не выявлены.

Оффлайн Arthwr

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Ещё одна новая работа по Монголии:

A dynamic 6,000-year genetic history of Eurasia's Eastern Steppe

Abstract
The Eastern Eurasian Steppe was home to historic empires of nomadic pastoralists, including the Xiongnu and the Mongols. However, little is known about the region's population history. Here we reveal its dynamic genetic history by analyzing new genome-wide data for 214 ancient individuals spanning 6,000 years. We identify a pastoralist expansion into Mongolia ca. 3000 BCE, and by the Late Bronze Age, Mongolian populations were biogeographically structured into three distinct groups, all practicing dairy pastoralism regardless of ancestry. The Xiongnu emerged from the mixing of these populations and those from surrounding regions. By comparison, the Mongols exhibit much higher Eastern Eurasian ancestry, resembling present-day Mongolic-speaking populations. Our results illuminate the complex interplay between genetic, sociopolitical, and cultural changes on the Eastern Steppe.

https://www.biorxiv.org/content/10.1101/2020.03.25.008078v1

Genetic group ID Time Period Affiliated cultures Number of Samples Sample IDs
eastMongolia_preBA pre-Bronze Age Unknown 1 SOU001
centralMongolia_preBA pre-Bronze Age Unknown 1 ERM001
Fofonovo_EN pre-Bronze Age Fofonovo  4 FNO001; FNO003; FNO006; FNO007
Afanasievo_Mongolia Early Bronze Age Afanasievo 2 SHT001; SHT002
Afanasievo_KUR001 Early Bronze Age Afanasievo 1 KUR001
Chemurchek Early Bronze Age Chemurchek 2 YAG001; IAG001
Chemurchek_KUM001 Early Bronze Age Chemurchek 1 KUM001
Altai_MLBA Middle Late Bronze Age Mönkhkairkhan, DSKC, Baitag 7 SBG001; BER002; ULI003; ULI001; BIL001; KHI001; ARS026*
KHU001 Middle Late Bronze Age Munkhkairkhan 1 KHU001
Ulaanzuukh_SlabGrave Middle Late Bronze Age; Early Iron Age Ulaanzuukh; Slab Grave 16 ULN010;ULN006;ULN009;ULN002;ULN005;BUL002;ULN001;ULN003;ULN007;BUL001;ULN015;DAR001;MIT001;BOR001;SHU001;PTO001
UAA001 Middle Late Bronze Age Munkhkairkhan 1 UAA001
ULZ001 Middle Late Bronze Age Unknown 1 ULZ001
UUS001 Middle Late Bronze Age DSKC 1 UUS001
TSI001 Middle Late Bronze Age Unknown 1 TSI001
ARS017 Middle Late Bronze Age Unknown 1 ARS017
Khovsgol_LBA Middle Late Bronze Age DSKC 17 ARS001,ARS002,ARS003,ARS005,ARS006,ARS007,ARS008,ARS009,ARS011,ARS012,ARS013,ARS014,ARS015,ARS016,ARS018,ARS024,ARS025
Chandman_IA Early Iron Age Uyuk 9 CHN007;CHN016;CHN012;CHN003;CHN008;CHN006;CHN001;CHN015;CHN004
earlyXiongnu_west early Xiongnu Xiongnu 6 SKT010;SKT001;SKT003;SKT009;SKT008;AST001
earlyXiongnu_rest early Xiongnu Xiongnu 6 SKT005;SKT012;SKT006;SKT002;SKT004;JAG001
SKT007 early Xiongnu Xiongnu 1 SKT007
lateXiongnu lateXiongnu Xiongnu 24 SAN001,UGU004,SOL001,DEL001,ULN004,BTO001,CHN010,DOL001,JAA001,KHO006,KHO007,TEV002,TUK003,UGU011,UVG001,TEV003,IMA001,IMA002,IMA003,IMA004,IMA005,IMA006,IMA007,IMA008
lateXiongnu_sarmatian lateXiongnu Xiongnu 13 TMI001,BUR001,NAI001,BUR002,BRL002,BUR003,BUR004,DUU001,HUD001,NAI002,UGU005,UGU006,UGU010
lateXiongnu_han lateXiongnu Xiongnu 8 ATS001,BAM001,BRU001,EME002,SON001,TUH001,TUH002,YUR001
YUR001 lateXiongnu Xiongnu 1 YUR001
TUK001 early Medieval Unknown 1 TUK001
earlyMed_Turk early Medieval Turk 7 ZAA004;TSB001;ULI002;NOM001;ZAA002;UGU001;ZAA007
earlyMed_Uigur early Medieval Uigur 12 OLN011;OLN001.B;OLN003;OLN005;OLN002;OLN008;OLN001.A;OLN010;OLN012;ZAA001;OLN004;OLN009
TUM001 early Medieval probably ancient Chinese 1 TUM001
OLN007 early Medieval Uigur 1 OLN007
lateMed_Khitan late Medieval Khitan 3 ZAA003;ZAA005;ULA001
lateMed_Mongol late Medieval Mongol 61 ARG001;ARG002;ARG003;KHV002;KNN001;CHD001;KRN001;KRN002;TSA001;TSA002;TSA003;MRI001;TSA005;BAY001;TAH002;NRC001;TAV001;TAV006;TAV005;DAR002;DAS001;BRG004;BRG005;KGK001;ZAR002;TSA004;KHN001;KHO001;BAZ001;GTO001;GUN002;BRG001;BRG002;IKU001;SHG002;SHG003;KHL001;UGO001;UGO002;UGU002;UGU003;RAH001;SHG001;ZAY001;KNU001;ZAM002;TSA007;SHR001;GAN002;TSA006;ERD001;TAV011;SHA001;ZAM001;ULN011;CHN014;DEE001;BAU001;BRL001;DUU002;UUS002
SHU002 late Medieval Unknown 1 SHU002

ARG001 M G2a1 R1a1a1b2a2b (R-Z2122)
ARG002 F M10a2 -
ARG003 M C4a2a1 C2b1a1b1 (C-F3830)
ARS001 M D4 Q1a2a1c (Q-L334; Q-L330)
ARS002 F D4j5 -
ARS003 M U5a2d1 N1c1a (N-M178)
ARS004 M F2a Q1a2 (Q-L942; Q-M346)
ARS005 M D4c1 Q1a2a1 (Q-L54)
ARS006 F C4a1a+195 -
ARS007 M A+152+16362 Q1a2a (Q-L475; Q-L53)
ARS008 M C4a2c Q1a2a1c (Q-L330)
ARS009 F D5a2a -
ARS011 M n/d Q1a2 (Q-L56; Q-M346)
ARS012 F C4a2a1 -
ARS013 F C -
ARS014 F C4a2c1 -
ARS015 M G3a Q1a1 (Q-Y706; Q-F1096)
ARS016 M A+152+16362+16189 Q1a2a1c (Q-L334; Q-L330)
ARS017 F G2a -
ARS018 M B5b1 Q1a (Q-M1117; Q-L472)
ARS024 F C4a1a1 -
ARS025 M D4b1a2a Q1a2a (Q-L475; Q-L53)
ARS026 M C4a1a+195 R1a1a1b2a2a (R-Z2123)
AST001 F I1a -
ATS001 F C4a1a -
BAA001 M D4j5a C2b (C-F2661; C-L1373)
BAM001 F D4j3 -
BAU001 F N1a1a1a1 -
BAY001 M D4e5b C2b1a1b (C-F3937; C-F1756)
BAZ001 F G3a3 -
BER002 M T1a1 N1c1 (N-L395; N-M46)
BIL001 F F1b1b -
BOR001 F - -
BRG001 F M7b1a1a2 -
BRG002 F G2a5 -
BRG004 M D4c2a CT (CT-M5613; CT-M168)
BRG005 M G1a N1c2b2 (N-L665)
BRL001 F G1a1a -
BRL002 F HV6 -
BRL003 M - C2b1a1b (C-F1756)
BRU001 M D5b1b2 J1a2b (J-P58)
BTO001 M Z4a Q1a2a1c (Q-L330)
BUL001 M M11b Q1a1a1 (Q-M120)
BUL002 M M10a1b G2a2b2a1a1c1a2a1a (G-FGC226; G-FGC249)
BUR001 F T2b -
BUR002 M U2e1 E1b1b1a1b2 (E-V22; E-L677)
BUR003 M J2b1a2a R1a1a1b (R-Z647; R-Z645)
BUR004 F G2a5 -
BUU001 F C4a1a4a -
CHD001 F F1g -
CHN001 M T1a1 R1a1a1b2a2b (R-Z2122)
CHN003 M U4a1 Q1a2a (Q-L213; Q-L53)
CHN004 F H91 -
CHN006 F U2e1h -
CHN007 M U2e1h Q1a2a1c1 (Q-L332; Q-L329)
CHN008 M F1b1f R1a1a1b2 (R-Z93)
CHN010 M D4j12 C2b1a1b (C-F1756)
CHN012 M T2b34 R1a1a1b (R-Z645)
CHN013 F D4g1b -
CHN014 F D4l2 -
CHN015 F N9a9 -
CHN016 M A8a Q1a2a1c1 (Q-L332; Q-L329)
DAR001 M F1b1f Q1b2 (Q-Y1151; Q-Y1150)
DAR002 F A11 -
DAS001 M N1a1a1a1a C2b1c (C-M546; C-F1918)
DEE001 F B4d1'2'3 -
DEK001 / SHR001 M R11a O2a2b1a1a3a (O-F474; O-F317)
DEL001 M D4j11 Q1a2a1c1 (Q-L332; Q-L329)
DOL001 F C4a1a -
DUU001 F D4b1a2a1 -
DUU002 M F2a R1a1a1 (R-Page7; R-M417)
EME002 F B5a2a1b -
EME003 M D5b1b2 O2a2b1a2a1 (O-F388; O-F46)
ERD001 F B4f -
ERM001 / ERM002 / ERM003 F A12 -
FNO001 M C4 N
FNO003 F C -
FNO006 F C4a1a3 -
FNO007 F F1b1+@152 -
GAN002 F A11a -
GTO001 F M7b1b -
GUN002 F F1a1 -
GUN003 M D5b1 N1 (N-L735)
HUD001 F K1a19 -
IAG001 M T2b2 R1b1a2 (R-CTS12478; R-M269)
IKU001 F F1b1b -
IMA001 F C4a1a+195 -
IMA002 F D4j5a -
IMA003 M D4j5a R1a1
IMA004 M G2a+152 Q1a2a
IMA005 M D4j5a N1c1a1a
IMA006 M A24 Q1a1a1
IMA007 F A+152+16362 -
IMA008 F N9a -
JAA001 F M10a1b -
JAG001 M F1b1f -
KGK001 F D4b1a2a1 -
KHI001 M R11 N1c2b2 (N-L665)
KHL001 F F1b1b -
KHN001 / KHN002 M D4j5a C2b1a1b1 (C-F3830)
KHO001 M D4 J (J-CTS10446; J-M304)
KHO003 F D4j5 -
KHO006 M C5b1b C2b1a1b (C-F3937; C-F1756)
KHO007 M - CT (CT-M5603; CT-M168)
KHU001 M C4a1a Q1a2a (Q-L55; Q-L53)
KHV001 M B4b1a3a CT (CT-M5606; CT-M168)
KHV002 M C5a1 N1c2b2 (N-L665)
KNN001 M Z3c O2a1c1a5 (O-M5420)
KNU001 F F1b1f -
KRN001 M Z3c C2b1b1 (C-M86)
KRN002 F A18 -
KUM001 F A -
KUR001 F A -
MIT001 M D1j Q1a1a1 (Q-M120)
MRI001 M D4m D (D-F1137; D-M174)
NAI001 M K1a12a1a J2a1h2 (J-L25)
NAI002 / NAI003 M C4a2c1 R1a1a1b1 (R-Z283)
NOM001 M U7a3a R1a1a1b2 (R-Z93)
NRC001 F A -
OLN001.A F D4 -
OLN001.B M B5b1a Q1a1 (Q-F1215; Q-F1096)
OLN002 F T1a1 -
OLN003 M T1a1 R1b1a2 (R-PF6505; R-M269)
OLN004 F H4a1a1a -
OLN005 M H6b R1b1a2a2 (R-Z2105; R-CTS1078)
OLN007 M D4 C2e1a1a (C-F3864; C-M407)
OLN008 F D4i -
OLN009 F J1b1b1 -
OLN010 M G3a3 R1b1a2a2 (R-Z2105; R-CTS1078)
OLN011 M B5b2a R1b1a2a2 (R-CTS1078)
OLN012 F D4j10 -
PTO001 F D4j+(16286) -
RAH001 F F1b1e -
SAN001 F F1b1e -
SBG001 M G2b2 Q1a2a1 (Q-L54)
SHA001 M D4a1 C2b1c (C-F3795; C-F1918)
SHG001 M M9a1a C2b1a1b1 (C-F3830)
SHG002 M D4o1 N1c1a1 (N-L708)
SHG003 M G2a2a C2b1c (C-F4002; C-F1918)
SHT001 M N1a1a1a1 R1b1a2a1a (R-P311)
SHT002 M U5a1 J1a (J-CTS5368)
SHU001 M G2b2a Q1a1a1 (Q-M120)
SHU002 M D4b2b R1a1a1b (R-Z645)
SKT001 F U5a1i -
SKT002 M G2a1e R1b (R-M343)
SKT003 F I1a -
SKT004 F U2e1a1 -
SKT005 M F1b1f R1b1 (R-M415; R-P25_1)
SKT006 M G2a1e R1 (R-P236; R-M173)
SKT007 M C4 Q1a2a1c (Q-L334; Q-L330)
SKT008 M H Q1a2a1c (Q-L334; Q-L330)
SKT009 M I1a R1a1a1b (R-Z647; R-Z645)
SKT010 F I1a -
SKT011 F D5a2a1 -
SKT012 M C4a1a4a J2a1h2 (J-L25)
SOL001 F I4a -
SON001 M D4 NO (NO-M2308; NO-M2313)
SOU001 F F1b1 -
TAH002 M B4b1b D1a2a (D-P47)
TAK001 F G2a1 -
TAV001 M M9a R1a1a1b (R-Z645)
TAV005 M D4j+16311 R1b1a2(R-PF6475,R-M269)
TAV006 F HV2a2 -
TAV007 M D4j3 R1b1a2a1a1 (R-M405)
TAV011 M C4b1 R1a1a1b2a2a (R-Z2123)
TEV002 F C5b -
TEV003 M N9a1 C2b1a1b(C-F1756)
TMI001 F J2b1a2a -
TSA001 M T1a1 C2b1a1b1 (C-F3830)
TSA002 M D4o2a C2b1 (C-F1788; C-F1699)
TSA003 M D4j3 C2b1c (C-F3939; C-F1918)
TSA004 M D4a1 D1a1a (D-N1)
TSA005 M Z1a N1c1 (N-L395; N-M46)
TSA006 F F1b1f -
TSA007 F M8a2'3 -
TSB001 M H14b J1a (J-CTS5368)
TSI001 F F1b1f -
TUH001 M B5a2a1a C2b1b1 (C-M86)
TUH002 M B5a2a1a J2a (J-M410)
TUK001/TAV008 F H2a2b -
TUK002 M B5b2a2 O2a1c1a5 (O-M5420)
TUK003 F J1d5 -
TUM001 M F1c1a1 N1c2 (N-L666)
UAA001 M H15b1 NO1 (NO-M214)
UGO001 F K2a5b -
UGO002 M U5a1a1 O2a2b1a1 (O-Page23)
UGU001 M D2 J2a (J-L559; J-M410)
UGU002 M B4a2b1 R (R-P224; R-M207)
UGU003 F U3a -
UGU004 F C4a1a4a -
UGU005 M U4a1 R1a1a1b2a2a (R-Z2123)
UGU006 M Z1a J1 (J-M267)
UGU010 M - R1a1a1b2a2a (R-Z2123)
UGU011 F HV6 -
ULA001 M Y1a J2a2 (J-L581)
ULI001 M U2e1 R1a1a1b2a2b (R-Z2122)
ULI002 M B4c1a2a J2a (J-L559; J-M410)
ULI003 F H6a1b2 -
ULN001 M D5a2a1b Q1a1a1 (Q-M265; Q-M120)
ULN002 F C4a1b -
ULN003 F C4a1a -
ULN004 F C5a1 -
ULN005 F D4 -
ULN006 F C4 -
ULN007 M D4b1a2 Q1a1 (Q-F1096)
ULN009 F C4a1a -
ULN010 F D5a2a -
ULN011 M A2a3 O2a2b1a1a (O-F42; O-F8)
ULN015 F D5a2a1b -
ULZ001 F D4b1a2 -
UUS001 M D5a2a J1a2b (J-P58)
UUS002 M X2d1 Q1a1a1 (Q-M265; Q-M120)
UVG001 F G2a2 -
YAG001 M T1a1 R1b1a2a2 (R-CTS1078)
YUR001 M D5a2a1 N1 (N-L735)
ZAA001 F A24 -
ZAA002 M C4a2a1 C2b1a1b1 (C-F3830)
ZAA003 F F2a -
ZAA004 M D4b1a2a1 C2b1a1b1 (C-F3830)
ZAA005 M D4c1b1 J2a (J-L212; J-M410)
ZAA007 F D4l2 -
ZAM001 M D4q R1a1a (R-L449; R-M512)
ZAM002 M B6a D1a1a (D-N1)
ZAR002 F F1 -
ZAY001 M M9a1b2 O2a2b1a1a (O-F8)

Оффлайн Arthwr

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  • Y-ДНК: R1b-PF7563
  • мтДНК: K1c1e
Afanasievo:

KUR001   F   A   -
SHT001   M   N1a1a1a1   R1b1a2a1a (R-P311)
SHT002   M   U5a1   J1a (J-CTS5368)

Chemurchek:

IAG001   M   T2b2   R1b1a2 (R-CTS12478; R-M269)
KUM001   F   A   -
YAG001   M   T1a1   R1b1a2a2 (R-CTS1078)

 

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